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Hemidactylus fasciatus GRAY, 1842

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesBanded Leaf-toed Gecko 
SynonymHemidactylus fasciatus GRAY 1842
Leiurus ornatus GRAY 1845: 157 (fide LOVERIDGE 1947)
Hemidactylus formosus HALLOWELL 1857: 148 (fide LOVERIDGE 1947)
Hemidactylus fasciatus — BOULENGER 1885: 124
Hemidactylus fasciatus — BROADLEY 1998
Hemidactylus fasciatus — WAGNER et al. 2014 
DistributionGuinea (Conakry), Liberia, Côte d’Ivoire, Ghana, Togo

Type locality: unknown (fide WERMUTH 1965) Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesHolotype: BMNH XXI.24.a
Syntypes: (3) ANSP 7451-53; Liberia [Hemidactylus formosus] 
CommentDescription. A fairly large species of Hemidactylus, with a maximum SVL of 95 mm, and a maximum total length of 172 mm. Like in the other species of the group, the body and limbs are moderate; the snout is slightly longer than the distance between the eye and the ear opening, the ear- opening is suboval and oblique; the upper surface of the body is covered with small granules, which are intermixed with small, round, and convex tubercle scales, forming more than 20 less regular longitudinal rows; digits are dis- tinctly webbed at the base, and have six to eight lamellae under the inner digits, and nine to ten under the median ones. Like other species of the group, excluding H. iturien- sis, it is morphologically characterized by a long row of femoral pores in males, which are interrupted in the mid- dle, resulting in a high number of femoral pores (16–20, vs. 8 in H. ituriensis) on each side. In coloration it is dis- tinct from its close relative H. kyaboboensis sp. n., by lack- ing the various pale tubercle spots and the broad pale fram- ing of the dark body crossbands, and by having the dark body crossbands more distinct to the pale interspaces (see Fig. 1). It is distinct from other species of the group by having a broad dark band between the eyes and the neck (vs. a narrow stripe in H. coalescens sp. n. and vs. lack- ing in H. ituriensis), and by having a pale upper lip (vs. dark in H. eniangii sp. n.) (Wagner et al. 2014).

Distribution: The species was considered to be widespread in West Africa until Wagner et al. 2014 split it up into several species. The distribution given here is the restricted range.

Habitat: forests, walls of houses

Relationships. Genetic data was presented by Leaché & Fujita (2010), who showed that this species includes all populations that cluster with those from the western Guinean rainforest with strong support in the Bayesian species delimitation model. This species is closely related to the other West African species described herein (H. kyaboboensis sp. n.), which is so far only known from forests within the Dahomey Gap. See tree in Wagner et al. 2014.

LEACHÉ & FUJITA 2010 described three new species related to H. fasciatus (H. coalescens, H. eniangii, H. kyaboboensis) but were “not aware of any morphological or ecological characteristics that differentiate these lineages”. The new species are only defined by DNA sequences. BAUER et al. (2010) considered these new species as nomina nuda. They were later (again) described by Wagner et al. 2014 with more details, although the new descriptions have some issues too. See the entries of these species for details. 
References
  • Bauer, Aaron M., Matthew LeBreton, Laurent Chirio, Ivan Ineich and Marcell T. Kouete 2006. New species of Hemidactylus (Squamata: Gekkonidae) from Cameroon. African Journal of Herpetology 55 (2):83-93. - get paper here
  • Bauer, Aaron M.; James F. Parham, Rafe M. Brown, Bryan L. Stuart, Lee Grismer, Theodore J. Papenfuss, Wolfgang Böhme, Jay M. Savage, Salvador Carranza, Jesse L. Grismer,, Philipp Wagner, Andreas Schmitz0, Natalia B. Ananjeva and Robert F. Inger 2010. Availability of new Bayesian-delimited gecko names and the importance of character-based species descriptions. Proc. R. Soc. B - doi:10.1098/rspb.2010.1330 - get paper here
  • BAUER,A.M.; S. TCHIBOZO, O.S.G. PAUWELS & G. LENGLET 2006. A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20 - get paper here
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  • Böhme, Wolfgang, Mark-Oliver Rödel, Christian Brede & Philipp Wagner 2011. The reptiles (Testudines, Squamata, Crocodylia) of the forested southeast of the Republic Guinea (Guinée forestière), with a country-wide checklist. Bonn zoological Bulletin 60 (1): 35-61 - get paper here
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  • Leaché, Adam D.; Mark-Oliver Rödel, Charles W. Linkem, Raul E. Diaz, Annika<br />Hillers, and Matthew K. Fujita 2006. Biodiversity in a forest island: reptiles and amphibians of the Togo Hills, Kyabobo National Park, Ghana. Amphibian & Reptile Conservation 4 (1): 22-45 - get paper here
  • Leaché, Adam D.; Matthew K. Fujita, Vladimir N. Minin, and Remco R. <br /> Bouckaert 2014. Species Delimitation using Genome-Wide SNP Data. Syst Biol 2014 63: 534-542 - get paper here
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  • Wagner, P., Leaché, A. & Fujita, M.K. 2014. Description of four new West African forest geckos of the Hemidactylus fasciatus Gray, 1842 complex, revealed by coalescent species delimitation. Bonn zoological Bulletin 63 (1): 1–14 - get paper here
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