Hemidactylus frenatus DUMÉRIL & BIBRON, 1836

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Higher Taxa	Gekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies
Common Names	E: Chichak, Common House Gecko, South Asian House Gecko, Bridled house gecko G: Gewöhnlicher Halbfingergecko, Asiatischer Hausgecko S: Tuqueque, tuteca, largarrabo, salamanquesa Chinese: 疣尾蜥虎
Synonym	Hemidactylus javanicus FITZINGER 1826 (nomen nudum) Hemidactylus frenatus H. BOIE in SCHLEGEL 1826: 235 (nomen nudum) Hemidactylus frenatus KUHL & VAN HASSELT in SCHLEGEL 1826:235 [nomen nudum] Hemi-dactylus frenatus KUHL & VAN HASSELT in SCHLEGEL 1826: 290 (nomen nudum) Hemidactylus frenatus DUMÉRIL & BIBRON 1836: 366 Hemidactylus (Pnoepus) Bojeri FITZINGER 1843 Hemidactylus vittatus GRAY 1845 Hemidactylus punctatus JERDON 1853 Hemidactylus fraenatus BLEEKER 1857 Hemidactylus inornatus HALLOWELL 1861 Hemidactylus pumilus HALLOWELL 1861: 502 Gecko caracal TYTLER 1865: 547 Gecko chaus TYTLER 1865: 547 Hemidactylus longiceps COPE 1869: 320 Hemidactylus hexaspis COPE 1869: 320 Hemidactylus papuensis [MACLEAY] 1877 Hemidactylus tristis SAUVAGE 1879: 49 Hemidactylus frenatus — BOULENGER 1885: 120 Hemidactylus nigriventris LIDTH DE JEUDE 1905 Hemidactylus bowringii — STEJNEGER 1907: 172 Hemidactylus frenatus — DE ROOIJ 1915: 28 Hemidactylus nigriventris — DE ROOIJ 1915: 31 Hemidactylus frenatus — TAYLOR 1922 Hemidactylus vandermeer-mohri BRONGERSMA 1928 Hemidactylus mabouia — BARBOUR & LOVERIDGE 1929 (partim) Hemidactylus okinawensis OKADA 1936 Hemidactylus vandermeermohri WERMUTH 1965 Hemidactylus auritus POEPPIG (in OBST) 1977 Pnoepus papuensis — WELLS & WELLINGTON 1985: 14 Pnoepus frenatus — WELLS & WELLINGTON 1985 Pnoepus bojeri — WELLS & WELLINGTON 1985 Pnoepus vittatus — WELLS & WELLINGTON 1985 Pnoepus punctatus — WELLS & WELLINGTON 1985 Pnoepus inornatus — WELLS & WELLINGTON 1985 Pnoepus pumilus — WELLS & WELLINGTON 1985 Pnoepus caracal — WELLS & WELLINGTON 1985 Hemidactylus frenatus — LANZA 1990 Hemidactylus frenatus — LINER 1994 Hemidactylus frenatus — GLAW & VENCES 1994: 277 Hemidactylus frenatus — MANTHEY & GROSSMANN 1997: 235 Hemidactylus frenatus — COX et al. 1998: 84 Hemidactylus frenatus — COGGER 2000: 246 Pnoepus frenatus — WELLS 2002 Hemidactylys cf. frenatus — ANDREONE et al. 2003 Hemidactylus vandermeermohri — TEYNIÉ et al. 2010 Hemidactylus cf. frenatus — JESTRZEMSKI et al. 2013 Hemidactylus frenatus — SPAWLS et al. 2018: 88 Hemidactylus frentatus — NEPALI & SINGH 2022 (in error)
Distribution	Worldwide in tropical and subtropical regions. Asia: Philippines (Palawan, Panay, Calamian Islands, Luzon, Mindoro, Bohol, Masbate, Cebu, Agusan del Sur, Camiguin Sur, Dinagat Island etc.), Japan (Ryukyu, Bonin, Amamioshima etc.), Palau, Singapore, Polynesia, Micronesia (Caroline Islands: Pohnpei, Mortlock Islands, Chuuk etc.), Melanesia, Solomon Islands [McCoy 2000], New Guinea, Timor-Leste, Taiwan, China (Hong Kong, Guangdong, Hainan, S Yunnan, Guangxi), India (Assam, Tamil Nadu, Andhra Pradesh, Maharashtra, Madhya Pradesh, Odisha, Mizoram, Kerala, Gujarat), Pakistan, Andaman Islands, Nicobar Islands, Bangladesh, Bhutan, Nepal, Sri Lanka, Maldives, Malaysia (incl. Pulau Tioman, Pulau Pangkor, Johor: Pulau Besar), Burma (= Myanmar), Vietnam (Binh Thuan etc.), Thailand, Cambodia, Indonesia (Sumatra, Borneo, Java, Bali, Lombok, Sulawesi, Ambon, Komodo, Flores, Halmahera, Morotai, Natuna islands), Nauru, Vanuatu, Northern Mariana Islands (Saipan) (Hileman et al. 2020) Africa: coastal Somalia (Lanza 1990), Kenya, Djibouti (Mazuch et al. 2016). Madagascar, Mauritius, Reunion, Rodrigues, Comoro Islands (Mayotte), Nossi Be = Nosy Bé (probably), Nosy Mitsio, Seychelles, Chagos Archipelago Australia (Cook Island, CKI, Northern Territory, Queensland, Western Australia, New South Wales), Western Samoa, New Caledonia, Fiji (introduced) Oceania: Tonga Islands Americas: Introduced to Mexico (Guanajuato, Yucatan, Jalisco, Guerrero, Oaxaca, San Luis Potosí, Sinaloa, Chiapas, Colima, Morelos, Baja California Sur, Campeche, Tabasco, Quintana Roo, Veracruz, Tamaulipas, Querétaro, Nayarit, Puebla, Hidalgo, Zacatecas, Sonora, Aguascalientes), Belize, Guatemala, Panama, Honduras [HR 31: 53; Isla de Utila: HR 32: 57], Costa Rica, Nicaragua, El Salvador (HR 33: 65); introduced to Venezuela, Ecuador, Colombia (incl. Valle del Cauca) Introduced to Mariana Islands (Guam) and New Caledonia. Introduced to USA (Hawaii, Florida) Introduced to Cuba Introduced to the Galapagos Islands Introduced to Puerto Rico (Sanchez 2018) Introduced to Mexico (Oaxaca) (McKay & Milenkaya 2020) Intorduced to St. Lucia (Griffing et al. 2022) Introduced to Aruba (Behm et al. 2022) Introduced to Antigua (Lindsey et al. 0223) fragilis: Somalia (LANZA 1983) Type locality: Java (designated by LOVERIDGE 1947)
Reproduction	oviparous
Types	Lectotype: MNHN-RA 5135A (1344), Java, presented RMNH (larger of two specimens under 5135), designated by Wells & Wellington (1985). Brygoo (1991) provided measurements for both 5135 and 5135A (the second specimen under the number 5135 as listed by Cogger et al. (1983), the presumed source for the information used by Wells & Wellington). Both have SVL 51mm, but MNHN 5135A has the longer tail (96mm total length vs 84mm total length), and hence 5135A is presumably the specimen that was intended to be designated the lectotype. Syntypes: MNHN-RA 0023, MNHN-RA 2300, MNHN-RA 2312, MNHN-RA 2312A-C, MNHN-RA 2313, MNHN-RA 2313A, MNHN-RA 2314, MNHN-RA 5135 and MNHN-RA 5135A. Holotype: MCZ 5730 [longiceps] Syntype: USNM 7429 [Hemidactylus inornatus]
Diagnosis	Diagnosis (partial): Vásquez-Restrepo 2018 provide illustrated comparisons between the Colombian Hemidactylus, H. frenatus, H. garnotii, and H. angulatus.
Comment	Distribution: Not listed for Honduras and Belize by KÖHLER (2000). DUMÉRIL & BIBRON (1836) give as type locality a whole range of places including South Africa, Mauritius, Madagascar, etc. In the Pacific H. frenatus and H. garnotii are sympatric only in Hawaii, the Marquesas, and New Caledonia. H. frenatus may have displaced H. garnotii in several instances (BAUER & VINDUM 1990). The arrival of H. frenatus in the Mascarene islands decimated a radiation of six species of Nactus geckos, three now being entirely extinct and the remainder limited to relict populations on the offshore islets of Mauritius (Arnold, 2000; Cole et al., 2005). South American and Galápagos haplotypes are identical to those from Hawaii and Papua New Guinea, suggesting a common Melanesian origin for both Hawaii and South America. Literature records suggest that H. frenatus arrived in Colombia around the mid-‘90s, dispersed south into Ecuador in less than five years, and arrived in the Galápagos about one decade later (Torres-Carvajal 2015). Original description in Amarasinghe et al. 2009. Synonymy: Sometimes Schlegel is cited as the describer, who mentioned the species as a communication from H. Boie (”par M. Boïé”), without any details, hence Schlegel’s name is a nomen nudum. However, the real authors are Duméril & Bibron, 1836 who ascribed the name to Schlegel and stated that this was the name under which the species was listed in the Leiden museum. This apparently is true, but Schlegel is not responsible for the description as published in D&B III:366 (M. Hoogmoed, pers. comm., 2 Sep 2014).
Etymology	The specific epithet is derived from the Latin word frenum, meaning "bridle or bit." The name pertains to the dusky streaks on the sides of the head between the eyes and the nostrils.
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External links	IUCN Red List National Center for Biotechnology Information http://www.calacademy.org/research/herpetology/myanmar/checklist_lizards… http://www.biodiversityexplorer.org/reptiles/squamata/scincidae… http://www.dnr.cornell.edu/edge/Field_guide_lizards.htm http://www.amonline.net.au/herpetology/research/pdf/gekkonidae.pdf https://www.facebook.com/groups/stshc/posts/236435001914439/ http://www.ifrance.com/reptiles-passion/classification/amphisbene… http://www.SriLankanReptiles.com/ http://snakes.mpsbb.info/lizard.html https://www.facebook.com/groups/stshc/posts/254166006808005/ http://www.morphobank.org/index.php?action=download_media&media_id=42140 http://www.jcu.edu.au/dept/Zoology/herp/trept/geck.html https://www.facebook.com/groups/stshc/posts/529732129251390/ http://www.wildherps.com/families/Gekkonidae.html http://www.online.de/home/laluenne/frena01.htm http://homepage3.nifty.com/japrep/gekko/othersgekko/text/ehooguro.htm http://forum.kingsnake.com/tax/messages/939.html http://wildherps.com/species/H.frenatus.html http://www.arod.com.au/arod/reptilia/Squamata/Gekkonidae/Hemidactylus… https://bioweb.bio/faunaweb/reptiliaweb/FichaEspecie/Hemidactylus… http://www.wildherps.com/species/L.lugubris.html https://www.facebook.com/groups/stshc/posts/366762518881686 https://www.facebook.com/groups/stshc/posts/313565994201339/ http://www.andamannicobarsnakes.com/Lycodoncapucinus.html http://www.ecologyasia.com/verts/snakes/common_house-snake.htm http://www.arod.com.au/arod/reptilia/Squamata/Colubridae/Lycodon/capucinus http://www.flickr.com/photos/73879198@N00/3719166585/ https://www.facebook.com/groups/stshc/posts/337282298496375/ Google images

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