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Hemidactylus granosus HEYDEN, 1827

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common Names 
SynonymHemidactylus granosus HEYDEN 1827: 17
Hemidactylus verrucosus – RÜPPELL 1845: 300 (part.)
Hemidactylus turcicus – BOETTGER 1893: 29 (part.)
Hemidactylus turcicus – ANDERSON 1898: 80 (part.)
Hemidactylus turcicus turcicus – LOVERIDGE 1947: 143 (part.)
Hemidactylus turcicus turcicus – MERTENS & WERMUTH 1960: 79 (part.)
Hemidactylus turcicus – SALVADOR 1981: 84 (part.)
Hemidactylus turcicus – BAHA EL DIN 2006: 66 (part.)
Hemidactylus turcicus turcicus – BAHA EL DIN 2005: 19 (part.)
Hemidactylus turcicus turcicus – MERTENS 1967: 55
Hemidactylus sp. 1 – MORAVEC et al. 2011: 24
Hemidactylus sp. 1 – CARRANZA & ARNOLD 2012: 17
Hemidactylus sp. 1 – ŠMÍD et al. 2013: 3
Hemidactylus granosus — ŠMÍD et al. 2013 
DistributionEgypt (Sinai), Saudi Arabia

Type locality: “Ägypten, Arabien, und Abyssinien”. Restricted to “Arabia petraea” = Sinai, Egypt by MERTENS 1967. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproduction 
TypesLectotype SMF 8723 designated by Mertens (1967) 
CommentSynonymy: Hemidactylus granosus HEYDEN 1827 has been resurrected from the synonymy of H. turcicus by SMID et al. 2013.

Diagnosis. Hemidactylus granosus is a member of the ‘Hemidactylus saba species group’ within the Arabian radiation of the Arid clade as evidenced by the mtDNA and nDNA analyses. The species has the following combination of molecular and morphological characters: (1) Uncorrected genetic distance from H. saba: 9.9–10.2% in 12S, 14.5–15.5% in cytb; from Hemidactylus sp. 4: 10.2–12.3% in 12S, 11.2–13.5% in cytb; (2) small size, SVL 39.0–53.2 mm in males, 40.6–53.3 mm in females; (3) rather elongated head, head length 24–28% of SVL, head width 68–86% of head length, head depth 33–47% of head length; (4) tail length 107–130% of SVL; (5) uppermost nasals separated by a small shield in 89% of specimens; (6) large anterior postmentals in wide mutual contact, and always in contact with the 1st and 2nd lower labial; (7) 9–11 upper labials; (8) 7–9 lower labials; (9) 14–15 longitudinal rows of enlarged, subtriangular, distinctly keeled dorsal tubercles; (10) 7–8 lamellae under the 1st toe and 10–13 under the 4th toe; (11) ca. 6–8 tail segments bearing 6 pointed tubercles; (12) 4–7 preanal pores in males forming a continuous row on the left and right side; (13) subcaudals enlarged; (14) in life, dorsum pale buff with dark brown spots tending to form transverse bands or X-shaped markings, dark horizontal stripe in prefrontal and temporal region, tail with ca. 10–13 dark brown transverse bands, venter white [from  ŠMÍD et al. 2013].

Distribution: see map in ŠMÍD et al. 2016: Fig. 3. 
References
  • Anderson, J. 1898. Zoology of Egypt. Vol. l. Reptilia and Batrachia. B. Quaritch, London
  • Baha el Din, S. 2006. A guide to the reptiles and amphibians of Egypt. The American University in Cairo Press, Cairo. xvi + 359 pp.
  • Baha El Din, Sherif M. 2005. An overview of Egyptian species of Hemidactylus (Gekkonidae), with the description of a new species from the high mountains of South Sinai. Zoology in the Middle East 34: 11-26 - get paper here
  • Boettger, Oskar 1893. Katalog der Reptilien-Sammlung im Museum der Senckenbergischen Naturforschenden Gesellschaft in Frankfurt am Main. I. Teil (Rhynchocephalen, Schildkröten, Krokodile, Eidechsen, Chamäleons). Gebrüder Knauer, Frankfurt a. M., x + 140 pp. - get paper here
  • Carranza, S., Arnold, E. Nicholas 2012. A review of the geckos of the genus Hemidactylus (Squamata: Gekkonidae) from Oman based on morphology, mitochondrial and nuclear data, with descriptions of eight new species. Zootaxa 3378: 1–95 - get paper here
  • Heyden, C. H. G. von. 1827. Reptilien. In Rüppell, E. Atlas zu Reise im nördlichen Afrika. l. Zoologie. H. L. Brönner, Frankfurt a. M., pp. 1-24 - get paper here
  • Loveridge,A. 1947. Revision of the African lizards of the family Gekkondiae. Bull. Mus. Comp. Zool. Harvard 98: 1-469 - get paper here
  • Mertens, R. 1967. Die herpetologische Sektion des Natur-Museums und Forschungs-Institutes Senckenberg in Frankfurt am Main nebst einem Verzeichnis ihrer Typen. Senckenbergiana Biologica 48: 1-106 - get paper here
  • Milto, Konstantin D. 2017. New Records of Reptiles on the Red Sea Coast, Egypt, with Notes on Zoogeography Russ. J. Herpetol. 24 (1): 11-21 - get paper here
  • Rösler, Herbert 2015. Bemerkungen über einige Geckos der Zoologischen Staatssammlung München. Gekkota, Suppl. (2): 3-54
  • Rüppell, E. 1845. Verzeichnis der in dem Museum der Senckenbergischen naturforschenden Gesellschaft aufgestellten Sammlungen. Dritte Abteilung: Amphibien. Mus. Senckenbergianum 3 (3): 293- 316
  • Šmíd J, Moravec J, Kratochvíl L, Gvoždík V, Nasher AK, Busais SM, Wilms T, Shobrak MY, Carranza S 2013. Two newly recognized species of Hemidactylus (Squamata, Gekkonidae) from the Arabian Peninsula and Sinai, Egypt. ZooKeys 355: 79–107 - get paper here
  • Šmíd, Jiří; Mohammed Shobrak; Thomas Wilms; Ulrich Joger & Salvador Carranza 2016. Endemic diversification in the mountains: genetic, morphological, and geographical differentiation of the Hemidactylus geckos in southwestern Arabia. Org Divers Evol, DOI 10.1007/s13127-016-0293-3
 
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