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Hemidactylus squamulatus TORNIER, 1897

IUCN Red List - Hemidactylus squamulatus - Least Concern, LC
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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
SubspeciesHemidactylus squamulatus floweri WERNER 1908
Hemidactylus squamulatus squamulatus TORNIER 1896 
Common NamesE: Tornier's Leaf-toed Gecko, Nyika Gecko 
SynonymHemidactylus squamulatus TORNIER 1897: 10
Hemidactylus bocagei TORNIER 1897: 12 (fide LOVERIDGE 1947 — non H. bocagii BOULENGER 1885)
Hemidactylus werneri TORNIER 1897 (nom. nov. pro H. bocagei)
Hemidactylus tornieri MOCQUARD 1902 (nom. nov. pro H. bocagei)
Hemidactylus alluaudi ANGEL 1923 (fide LOVERIDGE 1947)
Hemidactylus tropidolepis var. squamulatus — CALABRESI 1923: 153
Hemidactylus tropidolepis squamulatus — MERTENS 1938
Hemidactylus tropidolepis squamulatus — LOVERIDGE 1947
Hemidactylus squamulatus — WERMUTH 1965: 84
Hemidactylus squamulatus — KLUGE 1993
Hemidactylus squamulatus — LARGEN & SPAWLS 2010: 314
Hemidactylus squamulatus — SPAWLS et al. 2018: 96

Hemidactylus squamulatus squamulatus TORNIER 1896
Hemidactylus squamulatus TORNIER 1896: 10
Hemidactylus squamulatus squamulatus — LOVERIDGE 1957
Hemidactylus squamulatus squamulatus — BROADLEY & HOWELL 1991: 10
Hemidactylus squamulatus squamulatus — RÖSLER 2000: 87
Hemidactylus squamulatus squamulatus — RÖSLER 2015

Hemidactylus squamulatus floweri WERNER 1907
Hemidactylus floweri WERNER 1907: 478
Hemidactylus tropidolepis floweri — LOVERIDGE 1947: 112
Hemidactylus squamulatus floweri — WERMUTH 1965: 85
Hemidactylus squamulatus floweri — RÖSLER 2015
Hemidactylus floweri — SPAWLS et al. 2023 
Distributionsquamulatus: NE Kenya south to C Tanzania; Type locality: Kakoma, Uganda, Tanzania.

floweri: Republic of South Sudan (RSS) (”Blue Nile”); Type locality: “Anglo-Ägyptischer Sudan” = Republic of South Sudan (RSS)”  
Reproductionoviparous 
TypesHolotype: ZMB 13315, male
Holotype: not located fide De Lisle et al. 2018 [floweri] 
Diagnosis 
Comment 
References
  • Angel, F. 1923. Description de deux lézards nouveaux, des genres Hemidactylus et Mabuia, provenant d’Afrique Orientale (Missions Alluaud et Jeannel). Bulletin du Muséum National d’Histoire Naturelle, Paris, 29:490-492.
  • Bauer, A.M. & Günther,R. 1991. An annotated type catalogue of the geckos (Reptilia: Gekkonidae) in the Zoological Museum , Berlin. Mitt. Zool. Mus. Berlin 67: 279-310 - get paper here
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Calabresi, Enrica 1923. Anfibi e Rettili dell'Africa orientale raccolti durante le Spedizioni Franchetti e Zammarano. Atti Soc. Ital. Sci. Nat. (Milano), 62: 145-163. - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Lanza, B. 1990. Amphibians and reptiles of the Somali Democratic Republic: check list and biogeography. Biogeographia, 14: 407-465 [1988] - get paper here
  • Largen, M.J.; Spawls, S. 2006. Lizards of Ethiopia (Reptilia Sauria): an annotated checklist, bibliography, gazetteer and identification. Tropical Zoology 19 (1): 21-109 - get paper here
  • Largen, M.J.; Spawls, S. 2010. Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp.
  • Loveridge, A. 1920. Notes on East African lizards collected 1915-1919, with description of a new genus and species of skink and new subspecies of gecko. Proc. Zool. Soc. London 1920: 131-167 - get paper here
  • Loveridge, A. 1923. Notes on East African lizards collected 1920-1923 with the description of two new races of Agama lionotus Blgr. Proc. Zool. Soc. London 1923: 935-969 - get paper here
  • Loveridge, A. 1942. Scientific results of a fourth expedition to forested areas in east and central Africa. IV. Reptiles. Bull. Mus. Comp. Zool. Harvard 91: 237-373 - get paper here
  • Loveridge, A. 1947. Revision of the African lizards of the family Gekkondiae. Bull. Mus. Comp. Zool. Harvard 98: 1-469 - get paper here
  • Loveridge, A. 1957. Check list of the reptiles and amphibians of east Africa (Uganda, Kenya, Tanganyika, Zanzibar). Bull. Mus. Comp. Zool. Harvard 117 (2): 153-362 - get paper here
  • Malonza, Vincent; Beryl A. Bwong, Vincent Muchai 2011. Kitobo Forest of Kenya, a unique hotspot of herpetofaunal divers. Acta Herpetologica 6 (2): 149-160 - get paper here
  • Mertens, R. 1938. Über eine herpetologische Sammlung aus dem Gebiete des Njarasa-Grabens Ost-Afrikas. Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen 2 (1): 1-9 [no page numbers]
  • Mocquard, F. 1902. Sur des reptiles et batraciens de l’Afrique Orientale Anglaise, du Gabon et de la Guinée Francaise (Région de Kouroussa). Bulletin du Muséum d’Histoire Naturelle, Paris, 8: 404—417. - get paper here
  • Rösler, Herbert 2015. Bemerkungen über einige Geckos der Zoologischen Staatssammlung München. Gekkota, Suppl. (2): 3-54
  • Šmíd, Jiří; Salvador Carranza, Lukáš Kratochvíl, Václav Gvoždík, Abdul Karim Nasher, Jiří Moravec 2013. Out of Arabia: A Complex Biogeographic History of Multiple Vicariance and Dispersal Events in the Gecko Genus Hemidactylus (Reptilia: Gekkonidae). PLoS One 8(5): e64018 - get paper here
  • Spawls, S. & Rotich, D. 1997. An annotated checklist of the lizards of Kenya. J. East African Nat. Hist. 86: 61-83
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Spawls, Stephen; Tomáš Mazuch& Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • Tornier, G. 1897. Zur Faunistik Deutsch-Ost-Afrikas. 3. Reptilien und Amphibien. Archiv für Naturgeschichte 1:63—66 - get paper here
  • Tornier,G. 1897. Die Kriechthiere Deutsch-Ostafrikas. Beiträge zur Systematik und Descendenzlehre. D. Reimer, Berlin, 164 pp. - get paper here
  • Werner, F. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. Franz Werner's nach nach dem Ägyptischen Sudan und Norduganda. XII. die Reptilien und Amphibien. [Mabuia mongallensis, Leptodira attarensis]. Sitzungsber. Akad. Wiss. Wien 116 [1907]: 1823-1926 - get paper here
  • Zwanzig, Bernd-Michael & Wolfgang Grossmann 2013. Halbfingergeckos: Hemidactylus. Die verkannte Gattung. Terraria-Elaphe 2013 (2): 14-27 - get paper here
 
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