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Hemidactylus tenkatei LIDTH DE JEUDE, 1895

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymHemidactylus Tenkatei LIDTH DE JEUDE 1895: 121
Hemidactylus brookii subtriedroides ANNANDALE 1905
Hemidactylus brookii subtriedroides — RÖSLER & GLAW 2010
Hemidactylus tenkatei — RÖSLER & GLAW 2010
Hemidactylus tenkatei — MAHONY 2011
Hemidactylus tenkatei — O’SHEA et al. 2012
Hemidactylus cf. tenkatei — SANCHEZ et al. 2012
Hemidactylus tenkatei — RÖSLER 2016 
DistributionIndonesia (Pulau Roti); Timor-Leste, Guam

subtriedroides: N Myanmar (Burma); Type locality: “Tsagain, Upper Burma”

Type locality: Rotti  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesLectotype: RMNH 4353 (fide E. Dondorp, pers. comm., 28 Jan 2019)f
Lectotype: BMNH [subtriedroides] 
DiagnosisDiagnosis: “Die zwei vorliegenden Exemplare von H. tenkatei unterscheiden sich von der Originalbeschreibung in folgenden Pholidose- merkmalen: 17-18 versus 16 Rückentuberkelreihen, 5 versus 3-4 Subdigitallamellen unter der 1. Zehe und 4 versus 8 Tuberkel im 1. Schwanzwirtel (7 Schwanztuberkel im 2. Schwanzwirtel bei ZSM 222/1912/2). Hemidactylus tenkatei unterscheidet sich von H. b. brookii aus Nepal durch eine größere KRL (57,5 mm versus 52,0 mm), eine robustere Körperform, einen breiteren Schwanz, größere Rücken- und Schwanztuberkel, gekielten versus glatten Oberarmschuppen und weniger Präanofemoralporen (14 versus 19-22) sowie mehr glatte Schuppen zwischen den Präanofemoralporen (5 versus 2-3). Von H. b. brookii aus Myanmar unterscheidet sich H. tenkatei durch gekielte Oberarmschuppen und dort fehlende Tuberkel (“Limbs scaled above and below, some large tubercles on anterior and dorsal surface of forearm, more numerous enlarged tu- bercles on dorsal surface of thigh and crus.” Zug et al. 2007). Von H. parvimaculatus unterscheidet sich H. tenkatei durch weniger Präanofemoralporen (14 versus 24-31) sowie mehr glatte Schuppen zwischen den Präanofemoralporen (5 versus 2-4). Anhand der Originalbeschreibung von H. subtriedroides lässt sich H. tenkatei vorläufig lediglich durch die weniger stachelförmigen Schwanztuberkel und eine offenbar deutlichere Rückenzeichnung unterscheiden” [from RÖSLER & GLAW 2010]. 
CommentSynonymy: The types of H. subtriedroides were misidentified as H. maculatus by ANDERSON 1879. Hemidactylus subtriedroides Annandale was considered a synonym of H. tenkatei Lidth de Jeude by Mahony (2011), but Kathriner et al. 2014 consider Hemidactylus (brookii) subtriedroides as distinct from H. tenkatei and suggest to retain the former as valid species. Lajmi et al. (2016) used morphological and molecular data to recommend the synonymy of H. tenkatei with the older nomen, H. murrayi.

Distribution: The Pulau Roti, West Timor and “Australian” specimens of subtriedroides examined by Mahony 2011 are likely introductions rather than naturally occurring populations. 
EtymologyNamed after Dr. H. ten Kate, the collector of the types.

Etymology (subtriedroides): The specific epithet subtriedroides is derived from the species H. subtriedrus with the Greek suffix –oides, meaning “like” or “looks like”, from which it was considered in the original description to be most similar in appearance. 
References
  • Das, Abhijit 2016. Addition to the Herpetofauna of Royal Manas National Park, Bhutan, with six new country records. Herpetology Notes 9: 261-278 - get paper here
  • KATHRINER, ANDREW; MARK O’SHEA & HINRICH KAISER 2014. Re-examination of Hemidactylus tenkatei van Lidth de Jeude, 1895: Populations from Timor provide insight into the taxonomy of the H. brookii Gray, 1845 complex (Squamata: Gekkonidae). Zootaxa 3887 (5): 583–599 - get paper here
  • Lajmi, Aparna; Varad B. Giri & K. Praveen Karanth 2016. Molecular data in conjunction with morphology help resolve the Hemidactylus brookii complex (Squamata: Gekkonidae). Org Divers Evol, DOI 10.1007/s13127-016-0271-9 - get paper here
  • Lidth de Jeude, Th. W. van. 1895. Reptiles from Timor and the neighbouring islands. Notes from the Leyden Museum 16: 119-127 - get paper here
  • Mahony, Stephen 2011. Taxonomic revision of Hemidactylus brookii Gray: a re-examination of the type series and some Asian synonyms, and a discussion of the obscure species Hemidactylus subtriedrus Jerdon (Reptilia: Gekkonidae). Zootaxa 3042: 37–67 - get paper here
  • O’SHEA, Mark; Caitlin SANCHEZ, Andrew KATHRINER, Sven MECKE, Venancio LOPES CARVALHO, Agivedo VARELA RIBEIRO, Zito AFRANIO SOARES, Luis LEMOS DE ARAUJO and Hinrich KAISER 2015. Herpetological Diversity of Timor-Leste: Updates and a Review of Species Distributions. Asian Herpetological Research 6 (2): 73–131 - get paper here
  • O’SHEA, Mark; Caitlin SANCHEZ, Scott HEACOX, Andrew KATHRINER, Venancio LOPES CARVALHO, Agivedo VARELA RIBEIRO, Zito AFRANIO SOARES, Luis LEMOS DE ARAUJO and Hinrich KAISER 2012. First Update to Herpetofaunal Records from Timor-Leste. Asian Herpetological Research 3 (2): 114–126 - get paper here
  • Rösler H 2016. Bemerkungen über einige Geckoarten der Zoologischen Staatssammlung München. Gekkota Suppl. 3: 1-94
  • Rösler, H. & F. Glaw 2010. Morphologische Variation und Taxonomie von Hemidactylus brookii Gray, 1845 und Hemidactylus angulatus Hallowell, 1854 sowie phänotypisch ähnlicher Taxa. Spixiana 33 (1): 1-160 - get paper here
  • Sanchez, Caitlin; Venancio Lopes Carvalho,, Andrew Kathriner, Mark O’Shea and Hinrich Kaiser 2012. First report on the herpetofauna of the Oecusse District, an exclave of Timor-Leste. Herpetology Notes 5: 137-149
  • VAN EE, N., P. X. XIONG, D. V. YOUNG, K. L. KABAT, D. R. BRADKE, E. T. HILEMAN, S. W. GOTTE, D. G. MULCAHY, AND M. G. NAFUS 2020. Geographic Distribution: Hemidactylus tenkatei (Spotted House Gecko). USA: Guam: Tamuning. Herpetological Review 51: 540-541.
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
  • Zug, G.R. & Mulcahy, D.G. 2019. Identification guide Amphibians and reptiles of South Tanintharyi. Fauna & Flora International, 101 pp.
 
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