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Hemiphyllodactylus uga GRISMER, WOOD, THURA, ZIN, QUAH, MURDOCH, GRISMER, LI, KYAW & LWIN, 2018

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesE: Uga’s Slender Gecko 
SynonymHemiphyllodactylus uga GRISMER, WOOD, THURA, ZIN, QUAH, MURDOCH, GRISMER, LI, KYAW & LWIN 2018: 310
Hemphyllodactylus sp. nov. 8. — GRISMER et al. 2013: 872
Hemphyllodactylus sp. nov. 8. — GRISMER et al. 2014: 67
Hemphyllodactylus sp. nov. 8. — GRISMER et al. 2014: 490
Hemphyllodactylus sp. nov. 8. — NGO et al. 2014: 541
Hemphyllodactylus sp. nov. 8. — GRISMER et al. 2015: 861 
DistributionMyanmar (Mandalay)

Type locality: handrail of elevated timber walkway in Kandawgyi National Gardens, Pyin Oo Lwin, Mandalay Region, Myanmar (21.9931°N 96.4711°E; 1125 m in elevation  
Reproduction 
TypesHolotype: USNM 570733, adult male collected on 22 August 2003 at 1955 hrs by George R. Zug.
Paratypes. USNM 570732, Adult female collected on 14 August 2003 at ~1500 hrs by monastic brother in rockpile, Yangon Monastery, 11 km ENE of Pyin Oo Lwin, Mandalay Region, Myanmar (22°04.050 N 96° 33.811E, ~900 m elevation); adult females (USNM 570734, 570735) same locality as holotype, 23 August 2003 by San Lwin Oo ~1900 hrs and 20 August 2003 by Jeremy F. Jacobs from jaws of a spider ~1970 hrs, respectively. 
DiagnosisDiagnosis. Hemiphyllodactylus uga sp. nov. can be separated from all other species of Hemiphyllodactylus by possessing the unique combination of having a maximum SVL of 39.0 mm; 8–11 chin scales; enlarged postmentals; five circumnasal scales; two or three intersupranasals (=postrostrals); nine or 10 supralabials; 8–10 infralabials; 13–15 longitudinally arranged dorsal scales at midbody contained within one eye diameter and 6–8 ventral scales; varied digital formulae on hands and feet (Table 5); two or three subdigital lamellae on the first finger and toe; 18–22 continuous, pore-bearing, femoroprecloacal scales in males; no plate-like subcaudal scales; adult females not yellow; a dark postorbital stripe extending to base of neck variably present; variable pairs of paravertebral light spots on trunk; no dark, dorsolateral stripe; no dark paravertebral markings on trunk; dorsal body pattern usually with dark zig-zag markings; postsacral marking not bearing light-colored anteriorly projecting arms; and ceacum and gonads unpigmented. These characters are scored across all species of the eastern Myanmar clade in Tables 5 and 6 and from all other species of Hemiphyllodactylus from southern China and western Thailand (clades 3 and 4 in Grismer et al. [2017b:Table 3]).

Comparisons. The molecular analyses indicate that Hemiphyllodactylus uga sp. nov. is embedded within eastern Myanmar clade and forms a monophyletic group with H. linnwayensis and H. ywnaganensis sp. nov. See Comparisons section above for comparisons with H. ywanganensis. Hemiphyllodactlus uga sp. nov. is separated from H. montawaensis, H. tonywhitteni, and H. linnwayensis in have a higher mean number of chin scales (9.3 vs 6.3, p = 0.03; 9.3 vs 6.6, p = 0.05; 9.3 vs 5.0, p = 0.003, respectively; Table 7). It differs further from H. linnwayensis, H. tonywhitteni, and H. montawaensis in having a lower mean number of lamellae on the first toe (2.3 vs 4.5, p = 0.03; 2.3 vs 3.3 p = 0.002, respectively; Table 7). It differs even further from H. montawaensis in that adult females are not yellow and it lacks a reticulated dorsal pattern (Table 5). 
Comment 
EtymologyThe specific name recognizes and honors the late U Uga. He was a conservationist and a former director of the Nature and Wildlife Conservation Division (NWCD), Myanmar Forestry Department. He encouraged Joseph B. Slowinski and George R. Zug to do an all-country herpetofaunal survey and established the administrative protocol to establish and support survey teams of NWCD wildlife rangers. These teams working independently and with CAS and USNM collaborators were the essential factor for the high productivity and success of the Myanmar Herpetological Survey (MHS). 
References
  • GRISMER, L. LEE; AWAL RIYANTO, DJOKO T. ISKANDAR & JIMMY A. MCGUIRE 2014. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Pulau Enggano, southwestern Sumatra, Indonesia. Zootaxa 3821 (4): 485–495
  • Grismer, L. Lee; Perry L. Wood Jr, Shahrul Anuar, Evan S. H. Quah, Mohd Abdul Muin, Chan Kin Onn, Alexandra X. Sumarli and Ariel I. Loredo 2015. Repeated evolution of sympatric, palaeoendemic species in closely related, co-distributed lineages of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) across a sky-island archipelago in Peninsular Malaysia. Zoological Journal of the Linnean Society 174 (4): 859–876; DOI: 10.1111/zoj.12254 - get paper here
  • GRISMER, L. LEE; PERRY L. WOOD, JR., GEORGE R. ZUG, MYINT KYAW THURA, MARTA S. GRISMER, MATTHEW L. MURDOCH, EVAN S. H. QUAH & AUNG LIN 2018. Two more new species of Hemiphyllodactylus Bleeker (Squamata: Gekkonidae) from the Shan Hills of eastern Myanmar (Burma). Zootaxa 4483 (2): 295–316 - get paper here
  • GRISMER, L. LEE;PERRY L. WOOD, JR. & MICHAEL COTA 2014. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from northwestern Thailand. Zootaxa 3760 (1): 067–078 - get paper here
  • Grismer, L.L.; PERRY L. WOOD Jr, SHAHRUL ANUAR, MOHD ABDUL MUIN, EVAN S. H. QUAH, JIMMY A. MCGUIRE,, RAFE M. BROWN,0, NGO VAN TRI and PHAM HONG THAI 2013. Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia. Zoological Journal of the Linnean Society 169: 849–880
  • NGO VAN TRI; L. LEE GRISMER, PHAM HONG THAI & P. L. WOOD, JR. 2014. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Ba Na–Nui Chua Nature Reserve, Central Vietnam. Zootaxa 3760 (4): 539–552 - get paper here
 
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