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Hemiphyllodactylus ywanganensis GRISMER, WOOD, THURA, ZIN, QUAH, MURDOCH, GRISMER, LI, KYAW & LWIN, 2018

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesE: Ywangan Slender Gecko 
SynonymHemiphyllodactylus ywanganensis GRISMER, WOOD, ZUG, THURA, GRISMER, MURDOCH, QUAH & LIN 2018: 303 
DistributionMyanmar (Shan)

Type locality: 2.7 km southwest of Ywangan, Ywangan Township, Taunggyi District, Shan State, Myanmar (21.14643°N, 96.42178°E; 1157 m in elevation.  
Reproductionoviparous. The holotype is a gravid female carrying two eggs. The fact that the female was gravid indicates that March falls within the reproductive season of this species. 
TypesHolotype: LSUHC 13139, adult male collected on 28 March 2017 at 1600 hrs by L. Lee Grismer, Matthew L. Murdoch, and Toby Smith. Paratype. Adult female (LSUHC 13138), same collection data as the holotype. 
DiagnosisDiagnosis. Hemiphyllodactylus ywanganensis sp. nov. can be separated from all other species of Hemiphyllodactylus by possessing the unique combination of having a maximum SVL of 38.0 mm; 10 chin scales; enlarged postmentals; five circumnasals; two or three intersupranasals (=postrostrals); nine or 10 supralabials; 10 infralabials; 13–15 longitudinally arranged dorsal scales at midbody and seven or eight ventral scales contained within one eye diameter; varied digital formulae (Tables 5, 6, & 7); three subdigital lamellae on the first finger; two or three subdigital lamellae on the first toe; 26 continuous pore-bearing femoroprecloacal scales in the male; no plate-like subcaudal scales; adult females not being yellow; a dark postorbital stripe not extending onto trunk; no pairs of paravertebral light spots on trunk; no dark, diffuse, dorsolateral stripe; dark paravertebral markings on trunk; dorsal body pattern not unicolor; postsacral marking bearing light-colored anteriorly projecting arms; and unpigmented ceacum and gonads. These characters are scored across all species of the eastern Myanmar clade in Tables 5 and 6 and from all other species of Hemiphyllodactylus from southern China and western Thailand (clades 3 and 4 in Grismer et al. [2017:Table 3]).

Comparisons. The molecular analyses indicate that Hemiphyllodactylus ywanganensis sp. nov. is embedded within eastern Myanmar clade and forms a monophyletic group with H. linnwayensis and H. uga sp. nov. Although not statistically significant due to small sample sizes, it is separated from H. uga sp. nov. by having 26 (n = 1) as opposed to 18–22 (n = 2) femoroprecloacal pores in males. Additional samples of each will bear out the significance or further insignificance of this difference. It does differ from H. uga sp. nov. in having, as opposed to lacking dark, paravertebral markings on the dorsum and light-colored, anteriorly projecting arms of the postsacral marking and lacking, as opposed to variably having, pairs of paravertebral light spots on trunk. From H. linnwayensis, H. ywanganensis sp. nov. it differs further in lacking distinct pairs of light-colored paravertebral spots on the dorsum and having a significantly higher mean number of chin scales (10.0 vs 5.0, p = 0.003; Table 7). Similarly, it differs from H. tonywhitteni and H. montawaensis in having a significantly higher mean number of chin scales (10.0 vs 6.6, p = 0.04 and 10.0 vs 6.3, p = 0.03, respectively) and from H. montawaensis it differs even further in having more femoroprecloacal pores in males (26, n = 1 vs 19–21, n = 2) and a non-reticulated dorsal pattern. 
CommentHabitat: The area surrounding the collection site at the type locality is a generally flat, open and disturbed landscape with low, short, vegetated karstic ridges and small, isolated, scattered karst outcrops (Fig. 6). The holotype was found 1 m above the ground on the trunk of a small tree and the paratype was found approximately 10 m away 1 m above the ground on a karst boulder. 
EtymologyThe specific epithet, ywanganensis, is a noun in apposition in reference to the type locality being near the town of Ywangan, Shan State. 
References
  • GRISMER, L. LEE; PERRY L. WOOD, JR., GEORGE R. ZUG, MYINT KYAW THURA, MARTA S. GRISMER, MATTHEW L. MURDOCH, EVAN S. H. QUAH & AUNG LIN 2018. Two more new species of Hemiphyllodactylus Bleeker (Squamata: Gekkonidae) from the Shan Hills of eastern Myanmar (Burma). Zootaxa 4483 (2): 295–316 - get paper here
 
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