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Hemiphyllodactylus zugi NGUYEN, LEHMANN, LE DUC, DUONG, BONKOWSKI & ZIEGLER, 2013

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesE: Zug’s Slender Gecko
G: Zugs Halbblattfingergecko
Vietnamese: Thạch sùng dẹp zug 
SynonymHemiphyllodactylus zugi NGUYEN, LEHMANN, LE DUC, DUONG, BONKOWSKI & ZIEGLER 2013 
DistributionN Vietnam (Cao Bang), probably neighboring China (Guangxi)

Type locality: karst forest near Ban Nua Village (22o42.164’N, 106o38.317’E, at an elevation of 450 m a.s.l.), An Lac Commune, Ha Lang District, Cao Bang Province, northern Vietnam. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproduction 
TypesHolotype: IEBR A.2013.20 (CB 2012.85), adult male
Paratypes. Three adult specimens from karst forest near Ban Coong Village, Duc Quang Commune, Ha Lang District, Cao Bang Province, northern Vietnam: ZFMK 94781 (CB 2012.130), male, collected on 2 May 2012 by T. Lehmann, S. Herbst, and H. T. An near the entrance of a cave (22°42.768’N, 106°39.610’E, at an elevation of 601 m a.s.l.); IEBR A.2013.21 (CB 2012.137), male, collected on 3 May 2012 by T. Lehmann, S. Herbst, and H. T. An on a tree beside a forest path (22°42.864’N, 106°39.947’E, at an elevation of 435 m a.s.l.); and ZFMK 94782 (CB 2012.188), female, collected on 22 May 2012 by T. Q. Nguyen, H. T. An, T. Lehmann and S. Herbst on a rock boulder beside a forest path (22°43.202’N, 106°39.389’E, at an elevation of 448 m a.s.l.). 
CommentDiagnosis. The new species from northern Vietnam differs from the remaining congeners of the genus Hemiphyllodactylus by a combination of the following characters: a bisexual taxon; average SVL of adult males 41 mm, of adult female 46.6 mm; chin scales bordering mental and first infralabial distinctly enlarged; digital lamellae formulae 3-4-4-4 (forefoot) and 4-5-5-5 (hindfoot); femoral and precloacal pore series continuous, 18–21 in total in males, absent in female; cloacal spur single in males; dorsal trunk pattern of dark brown irregular transverse bands; dark lateral head stripe indistinct; upper zone of flank with a series of large light spots, edged above and below in dark grey; caecum and gonadal ducts unpigmented.

Behavior: nocturnal 
EtymologyNamed in honour of Professor Dr. George Zug, National Museum of Natural History, Smithsonian Institution (Washington DC, USA), in recognition of his contribution to herpetological research in Southeast Asia, particularly the genus Hemiphyllodactylus. 
References
  • GRISMER, L. LEE; AWAL RIYANTO, DJOKO T. ISKANDAR & JIMMY A. MCGUIRE 2014. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Pulau Enggano, southwestern Sumatra, Indonesia. Zootaxa 3821 (4): 485–495
  • Grismer, L. Lee; Perry L. Wood Jr, Shahrul Anuar, Evan S. H. Quah, Mohd Abdul Muin, Chan Kin Onn, Alexandra X. Sumarli and Ariel I. Loredo 1860. Repeated evolution of sympatric, palaeoendemic species in closely related, co-distributed lineages of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) across a sky-island archipelago in Peninsular Malaysia. Zoological Journal of the Linnean Society 174 (4): 859–876; DOI: 10.1111/zoj.12254 - get paper here
  • GUO, WEIBO; KAIYA ZHOU, JIE YAN & PENG LI 2015. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from western Yunnan, China. Zootaxa 3974 (3): 377–390
  • NGUYEN, TRUONG QUANG; TANJA LEHMANN, MINH DUC LE, HA THUY DUONG, MICHAEL BONKOWSKI & THOMAS ZIEGLER 2013. A new species of Hemiphyllodactylus (Reptilia: Gekkonidae) from northern Vietnam. Zootaxa 3736 (1): 089–098 - get paper here
 
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