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Herpetoreas sieboldii GÜNTHER, 1860

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Siebold’s Keelback, Sikkim Keelback 
SynonymHerpetoreas sieboldii GÜNTHER 1860: 156
Herpetoreas sieboldii — GÜNTHER 1864: 257
Herpetoreas sieboldii — THEOBALD 1868: 48
Herpetoreas sieboldii — THEOBALD 1876: 172
Amphiesma sieboldii — DAS 1996: 53
Amphiesma sieboldii — DAVID et al. 2005
Amphiesma sieboldii — WALLACH et al. 2014: 33
Herpetoreas sieboldii — GUO et al. 2014
Herpetoreas sieboldii — LALREMSANGA et al. 2022 
DistributionPakistan, India (Uttar Pradesh, Sikkim, Punjab), Nepal, Myanmar (= Burma), Bangladesh, Bhutan

Type locality: "Sikkim, Himalaya (7500 feet above the level  
Reproductionoviparous 
TypesHolotype: BMNH 1946.1.13.16, a 917 mm male (Hermann, Adolphe and R. von Schlagintweit, 1854-1858). 
DiagnosisDiagnosis (genus). Semi-aquatic snakes with keeled dorsal scales. The posterior maxillary tooth longest, in a continuous series with the anterior ones. Body and tail slender, compressed. Two nasals, one loreal, one anterior, two posterior oculars. Scales moderately elongate, keeled, in nineteen rows. Eye is moderate size (Günther, 1860) [from Guo et al. 2014]. 
CommentSynonymy: Has been synonymized with A. platyceps. Both species are very similar and characters are broadly overlapping (see also TILLACK 2003).

Type species: Herpetoreas sieboldii GÜNTHER 1860: 156 is the type species of the genus Herpetoreas GÜNTHER 1860.

For a comparison of characters across members of Herpetoreas see Das et al. 2020: 313 (Table 3) and Lalremsanga et al. 2022 (Table 2).

Key to the species of Herpetoreas: Lalremsanga et al. 2022 
References
  • Das, A., Gower, D. J., & Deepak, V. 2020. Lost and found: rediscovery and systematics of the Northeast Indian snake Hebius pealii (Sclater, 1891). Vertebrate Zoology 70 (3): 305–318 - get paper here
  • Das, I. 2012. A Naturalist's Guide to the Snakes of South-East Asia: Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. Oxford J, ohn Beaufoy Publishing - get paper here
  • David, P.; Vogel, G. & Pauwels, O.S.G. 2005. On the occurrence of Amphiesma bitaeniatum (WALL, 1925) in Vietnam, with preliminary remarks on the group of Amphiesma parallelum (BOULENGER, 1890) (Serpentes, Colubridae, Natriciniae). Salamandra 41 (4): 167-178 - get paper here
  • DAVID, PATRICK; ISHAN AGARWAL, RAMANA ATHREYA, ROSAMMA MATHEW,<br />GERNOT VOGEL & VIRAL K. MISTRY 2015. Revalidation of Natrix clerki Wall, 1925, an overlooked species in the genus Amphiesma Duméril, Bibron & Duméril, 1854 (Squamata: Natricidae). Zootaxa 3919 (2): 375–395 - get paper here
  • Günther, A. 1860. Contributions to a knowledge of the reptiles of the Himalaya mountains. - I. Descriptions of the new species. II. List of Himalayan reptiles, with remarks on their horizontal distribution. Proc. Zool. Soc. London 1860: 148-175 - get paper here
  • Günther, A. 1860. Reptiles of Siam. Proc. Zool. Soc. London 1860: 113-117 - get paper here
  • Günther, A. 1864. The Reptiles of British India. London (Taylor & Francis), xxvii + 452 pp. - get paper here
  • GUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440 - get paper here
  • Khan, M.S. 2002. Die Schlangen Pakistans [English edition as “A Guide to the snakes of Pakistan”]. Edition Chimaira (Frankfurt am Main), 265 pp. [review in HR 34: 400, Russ J Herp 12: 79] - get paper here
  • Kramer, E. 1977. Zur Schlangenfauna Nepals. Revue Suisse de Zoologie 84 (3): 721-761. - get paper here
  • Lalremsanga, H. T., Bal, A. K., Vogel, G., & Biakzuala, L. 2022. Molecular phylogenetic analyses of lesser known colubrid snakes reveal a new species of Herpetoreas (Squamata: Colubridae: Natricinae), and new insights into the systematics of Gongylosoma scriptum and its allies from northeastern India. SALAMANDRA, 58(2): 101-115 - get paper here
  • Malnate, E. V. 1966. Amphiesma platyceps (BLYTH) and Amphiesma sieboldii (GÜNTHER): sibling species (Reptilia: Serpentes). J. Bombay Nat. Hist. Soc. 63 (1): 1-17 - get paper here
  • Theobald, WILLIAM 1868. Catalogue of reptiles in the Museum of the Asiatic Society of Bengal. J. Asiatic Soc. Bengal, Calcutta, 37 (extra number 146): (2), vi, 7-88 - get paper here
  • Theobald,W. 1876. Descriptive catalogue of the reptiles of British India. Thacker, Spink & Co., Calcutta: xiii + 238 pp. - get paper here
  • Tillack, F. 2003. Über die Verbreitung und Biologie der Himalaya-Gebirgswassernatter Amphiesma platyceps (BLYTH 1854) und einen Fall von Amphigonia retardata (Serpentes: Colubridae: Natricinae). Sauria 25 (1): 21-27 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wangyal, Jigme Tshelthrim 2019. The status of herpetofauna of Bhutan. District Forest Office, District Administration, Trashigang, Bhutan, 20-39 - get paper here
  • Ziegler, T., and Quyet, L.K. 2006. A new natricine snake of the genus Amphiesma (Squamata: Colubridae: Natricinae) from the central Truong Son, Vietnam. Zootaxa 1225: 39-56 - get paper here
 
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