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Heterodactylus imbricatus SPIX, 1825

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Higher TaxaGymnophthalmidae (Gymnophthalminae), Heterodactylini, Sauria, Gymnophthalmoidea, Squamata (lizards) 
Subspecies 
Common NamesRio de Janeiro Teiid 
SynonymHeterodactylus imbricatus SPIX 1825
Chirocolus imbricatus — WAGLER 1830
Chirocolus imbricatus — SCHINZ 1833
Heterodactylus imbricatus — DUMÉRIL & BIBRON 1839: 447
Chirocolus imbricatus — GRAY 1845
Chirocolus imbricatus — FITZINGER 1867
Heterodactylus imbricatus — PETERS & DONOSO-BARROS 1970: 144
Heterodactylus imbricatus — CASTOE et al. 2004 
DistributionBrazil (Rio de Janeiro, Minas Gerais, São Paulo, Espírito Santo)

Type locality: Estado Rio de Janeiro, Brazil Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesHolotype: ZSM 108/0 (lost) 
CommentType species: Heterodactylus imbricatus SPIX 1825 is the type species of the genus Heterodactylus SPIX 1825.

Tribus: Goicoechea et al. 2016 erected a new tribe, Heterodactylini, for the genera Caparaonia, Colobodactylus, and Heterodactylus.

Synonymy: Because the type species of Chirocolus (H. imbricatus) is also the type species of Heterodactylus, Chirocolus is a junior objective synonym of Heterodactylus. Even though the genus name Heterodactylus is older than Chirocolus, Chirocolini has priority over Heterodactylini (ICZN Article 40.1); therefore, the valid family-group name for the clade including Heterodactylus is Chirocolini Gray 1838. Colli et al. 2015 defined the tribe Chirocolini to include Caparaonia, Colobodactylus, Heterodactylus. 
References
  • Bruscagin, R. T., M. Dixo, S. Famelli & J. Bertoluci 2017. Patch size effects on richness, abundance, and diversity of leaf-litter lizards from Atlantic rainforest fragments.  Salamandra 53 (1): 59-65 - get paper here
  • Castoe, T.A.; Doan, T.M. & Parkinson, C.L. 2004. Data partitions and complex models in Bayesian analysis: the phylogeny of Gymnophthalmid lizards. Systematic Biology 53 (3): 448-469 - get paper here
  • COLLI, GUARINO R.; MARINUS S. HOOGMOED, DAVID C. CANNATELLA, JOSÉ CASSIMIRO, JERRIANE OLIVEIRA GOMES, JOSÉ MÁRIO GHELLERE, PEDRO M. SALES NUNES, KÁTIA C. M. PELLEGRINO, PATRICIA SALERNO, SERGIO MARQUES DE SOUZA & MIGUEL TREFAUT RODRIGUES 2015. Description and phylogenetic relationships of a new genus and two new species of lizards from Brazilian Amazonia, with nomenclatural comments on the taxonomy of Gymnophthalmidae (Reptilia: Squamata) Zootaxa 4000 (4): 401–427 - get paper here
  • Duméril, A. M. C. and G. Bibron. 1839. Erpétologie Générale on Histoire Naturelle Complète des Reptiles. Vol.5. Roret/Fain et Thunot, Paris, 871 pp. - get paper here
  • Goicoechea, N., Frost, D. R., De la Riva, I., Pellegrino, K. C. M., Sites, J., Rodrigues, M. T. and Padial, J. M. 2016. Molecular systematics of teioid lizards (Teioidea/Gymnophthalmoidea: Squamata) based on the analysis of 48 loci under tree-alignment and similarity-alignment. Cladistics, doi: 10.1111/cla.12150 - get paper here
  • Novelli, I.A.; P.S. Lucas; R.C. Santos. 2011. Reptilia, Squamata, Gymnophthalmidae, Heterodactylus imbricatus Spix, 1825: Filling gaps in the state of Minas Gerais. Check List 7 (1):30-31 - get paper here
  • ROCHA, CARLOS FREDERICO D.; HELENA G.BERGALLO, JOSÉ P. POMBAL JR., LENA GEISE, MONIQUE VAN SLUYS, RONALDO FERNANDES, ULISSES CARAMASCHI 2004. FAUNA DE ANFÍBIOS, RÉPTEIS E MAMÍFEROS DO ESTADO DO RIO DE JANEIRO, SUDESTE DO BRASIL Publ. Avul. Mus. Nac., Rio de Janeiro (104): 3-23 - get paper here
  • Rodrigues, Miguel Trefaut; Katia Cristina Machado Pellegrino, Marianna Dixo, Vanessa Kruth Verdade, Dante Pavan, Antônio Jorge Suzart Argolo, and Jack W. Sites Jr. 2007. A new genus of microteiid lizard from the Atlantic forests of State of Bahia, Brazil, with a new generic name for Colobosaura mentalis, and a discussion of relationships among the Heterodactylini (Squamata, Gymnophthalmidae). American Museum novitates (3565): 1-27 - get paper here
  • Silva-Lucas, Priscila; Iara Alves Novelli, Bernadete Maria de Sousa 2016. Assemblage of squamate reptiles in a natural remnant of Cerrado in southern Minas Gerais, Brazil Check List 12 (2): 1-9 - get paper here
  • Spix, J. B. von 1825. Animalia nova sive species nova lacertarum quas in itinere per Brasiliam annis MDCCCXVII-MDCCCXX jussu et auspicius Maximiliani Josephi I Bavariae Regis suscepto collegit et descripsit Dr. J. B. de Spix. Lipsiae: T. O. Weigel; F. S. Hübschmanni, Monachii, 26 pp.
 
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