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Letheobia sudanensis (SCHMIDT, 1923)

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Higher TaxaTyphlopidae (Afrotyphlopinae), Typhlopoidea, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesGaramba gracile blind-snake, Sudan Beaked Snake 
SynonymTyphlops sudanensis SCHMIDT 1923: 51
Typhlops sudanensis — ESSEX 1928
Rhinotyphlops sudanensis - ROUX-ESTÈVE 1974: 233
Typhlops sudanensis — PITMAN 1974
Rhinotyphlops sudanensis — MCDIARMID, CAMPBELL & TOURÉ 1999: 87
Letheobia sudanensis — BROADLEY & WALLACH 2007
Letheobia sudanensis — HEDGES et al. 2014 
DistributionZaire (Orientale), N Democratic Republic of the Congo, elevation 700–1300 m, Uganda

Type locality: Faradje (contra Schmidt 1923: 4, who erroneously lists the type locality as Garamba), Sudan [= Ituri, Democratic Republic of Congo] (03°43’N, 29°42’E, elevation 820 m). Map legend:
Type locality - Type locality.
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesHolotype: AMNH 11677; collected by H. Lang and J. P. Chapin, November 1911 
CommentDescription: Snout with an angular horizontal edge. Rostral very broad, truncated posteriorly; frontal crescentic, in contact with nasals; supraocular transverse, its lateral apex wedged between a very large nasal and a deep ocular, which in turn is wedged between the preocular and subocular; eye not visible; nasal suture arising from second labial; SIP X (N1/N2, P, P, S), one paratype (AMNH 11680) with SIP II (N2, P, P, S) and another paratype (MCZ 1599) with an aberrant SIP III-S (N2, P, P, S) on one side of head; scale rows 26-24-24 (rarely 24-24-24); MD 569–660; vertebrae 66–412; MD/V ratio 1.49–1.62; L/D ratio 57–95. Colourless. For abbreviations see L. caeca. From BROADLEY & WALLACH 2007.

Habitat: Mosaic of lowland rain forest and secondary grassland (Sudanese savanna). 
References
  • Broadley, Donald G. & Wallach, V. 2007. A review of East and Central African species of Letheobia Cope, revived from the synonymy of Rhinotyphlops Fitzinger, with descriptions of five new species (Serpentes: Typhlopidae). Zootaxa 1515: 31–68 - get paper here
  • Chifundera, K. 1990. Snakes of Zaire and their bites. Afr. Stud. Monogr. (Kyoto) 10(3): 137-157.
  • Essex, R. 1929. A note on three burrowing reptiles. Ann. Mag. nat. Hist. (10) 1: 268-270
  • Hedges, S.B., Marion, A.B., Lipp, K.M., Marin, J. & Vidal, N. 2014. A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology 49: 1–61 - get paper here
  • McDiarmid, R.W.; Campbell, J.A. & Touré,T.A. 1999. Snake species of the world. Vol. 1. Herpetologists’ League, 511 pp.
  • Pitman,C.R.S. 1974. A guide to the snakes of Uganda. Codicote, Wheldon & Wesley, L., 290 pp.
  • Roux-Estève, R. 1974. Révision systématique des Typhlopidae d'Afrique. Reptilia. Serpentes. Mém. nation. Hist. nat., Paris, (sér.A.) 87: 1-313
  • Schmidt, K. P. 1923. Contributions to the herpetology of the Belgian Congo based on the collection of the American Museum Congo Expedition, 1909-1915. Part II. Snakes, with field notes by Herbert Lang and James P. Chapin. Bull. Amer. Mus. nat. Hist. 49 (1): 1-146
 
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