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Liasis mackloti DUMÉRIL & BIBRON, 1844

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Higher TaxaPythonidae, Henophidia, Pythonoidea, Alethinophidia, Serpentes, Squamata (snakes)
SubspeciesLiasis mackloti mackloti DUMÉRIL & BIBRON 1844
Liasis mackloti dunni STULL 1932
Liasis mackloti savuensis BRONGERSMA 1956 
Common NamesE: Freckled Python, Macklot's Python
G: Macklots Python, Timor-Wasserpython 
SynonymLiasis Mackloti DUMÉRIL & BIBRON 1844: 440
Python timorensis MÜLLER 1844
Python timoriensis — MÜLLER 1857
Liasis corwallisius GÜNTHER 1879
Nardoa crassa MACLEAY 1885
Liasis mackloti — BOULENGER 1893: 79
Liasis mackloti — DE ROOIJ 1917: 17
Liasis fuscus PETERS 1873 (fide KLUGE 1993)
Morelia mackloti — UNDERWOOD & STIMSON 1990: 603
Liasis mackloti — MCDIARMID, CAMPBELL & TOURÉ 1999: 169
Katrinus mackloti — HOSER 2001
Liasis mackloti — ESCORIZA BOI 2005
Liasis mackloti mackloti — DE LANG 2011
Liasis mackloti — REYNOLDS et al. 2014
Liasis mackloti — WALLACH et al. 2014: 383

Liasis mackloti dunni STULL 1932
Liasis dunni — WALLS 1998
Katrinus mackloti dunni — HOSER 2001
Liasis mackloti dunni — RAWLINGS et al. 2004
Liasis dunni — CARMICHAEL 2007
Liasis mackloti dunni — DE LANG 2011
Liasis dunni — BARKER et al. 2015

Liasis mackloti savuensis BRONGERSMA 1956
Liasis savuensis — WALLS 1998
Katrinus savuensis — HOSER 2000
Liasis mackloti savuensis — RAWLINGS et al. 2004
Liasis savuensis — VIDAL et al. 2007
Liasis mackloti savuensis — HENDERSON & POWELL 2007
Liasis mackloti savuensis — DE LANG 2011
Liasis savuensis — BARKER et al. 2015 
DistributionIndonesia (Savu = Sawoe, Samoa, Alor, Roti, Semau = Samao, Timor, Wetar, Irian Jaya), Timor-Leste,
Papua New Guinea

Type locality: Timor

dunni: Wetar Island

savuensis: Sawu Island  
Reproductionoviparous 
TypesLectotype: MNHN-RA 1625 (BRONGERSMA 1968: 57)
Holotype: AMNH 32263 [dunni]
Holotype: BMNH 1897.6.21.33 (Stimson, 1969: 25), incorrectly stated as BMNH 1896.6.21.33 by McDowell (1975: 35); other syntypes: BMNH 1897.6.21.33-37 [savuensis] 
Diagnosis 
CommentType species: Liasis Mackloti DUMÉRIL & BIBRON 1844 is the type species of the genus Liasis GRAY 1842: 44 (fide Stimson and McDowell's 1986 request for conservation of Liasis), otherwise the type species is Liasis childreni, see Williams 1987: 69).

Synonymy: Kaiser et al. 2013 considered the generic name Katrinus Hoser 2000 invalid and rejected its use instead of Liasis. See comment on Katrinus in Liasis fuscus.

Hybridization: BANKS & SCHWANER (1984) describe a hybrid between Morelia spilota and Liasis mackloti.

Distribution: reports from coastal N Australia may be misidentifications of other Liasis species. 
EtymologyNamed after Heinrich Christian Macklot (1799-1832), a taxidermist who was appointed to assist members of the Dutch Natural Science Commission. 
References
  • Banks, C., Schwaner, T.D., 1984. Two cases of interspecific hybridization among captive Australian boid snakes [Python spilotes, Python amethistinus, Liasis mackloti.]. Zoo Biology 3: 221-227 - get paper here
  • Barker, D. G., Barker, T. M., Davis, M. A. and Schuett, G. W. 2015. A review of the systematics and taxonomy of Pythonidae: an ancient serpent lineage. Zoological Journal of the Linnean Society 175 (1): 1-19; doi: 10.1111/zoj.12267 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Brongersma, L.D. 1956. On two species of boid snakes from the Lesser Sunda Islands. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, Amsterdam 59C(3): 290–300
  • Brongersma,L.D. 1968. Proposal to Suppress Python timorensis MÜLLER, 1844, and Python timoriensis MÜLLER, 1857, and to Add Python timoriensis (PETERS, 1876) to the Official List (Reptilia, Serpentes, Boidae). Z. N. (S.) 1834. Bull. zool. Nomenclature 25: 55-59. - get paper here
  • Bulian, J. & W. Bröer 1987. Ein seltener Python und seine Nachzucht: Liasis mackloti. Litteratura Serpentium 7 (2): 95-96 - get paper here
  • Bulian, Joachim 1994. Ten years of breeding Liasis mackloti mackloti. Litteratura Serpentium 14 (2): 46-56 - get paper here
  • Bulian,J. & Broer, W. 1984. Ein seltener Python und seine Nachzucht: Liasis mackloti (DUMÉRIL & BIBRON 1844) (Serpentes: Boidae). Salamandra 20 (4): 205-211 - get paper here
  • Carmichael, C.K. 2007. PHYLOGEOGRAPHY OF THE INDONESIAN WATER PYTHON, LIASIS MACKLOTI SSP. (SQUAMATA: BOIDAE: PYTHONINAE): A COMPARATIVE APPROACH TOWARD RESOLVING PHYLOGENY. PhD Dissertation, U of S Mississippi, 153 pp.
  • de Rooij, N. DE 1917. The Reptiles of the Indo-Australian Archipelago. Il. Ophidia. Leiden (E. J. Brill), xiv + 334 S. - get paper here
  • Duméril, A. M. C. and G. Bibron. 1844. Erpetologie Générale ou Histoire Naturelle Complete des Reptiles. Vol.6. Libr. Encyclopédique Roret, Paris, 609 pp. - get paper here
  • Escoriza Boj, D. 2005. Australia. Reptiles and Amphibians, Part 2: Desert and tropical savanna. Reptilia (GB) (41): 52-57 - get paper here
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  • Hoser, Raymond 2000. A Revision of the Australasian pythons. Ophidia Review 1: 7-27
  • Jan, G. 1864. Iconographie générale des ophidiens. 7. Livraison. J.B. Bailière et Fils, Paris - get paper here
  • Kaiser, H.; Crother, B.I.; Kelly, C.M.R.; Luiselli, L.; O’Shea, M.; Ota, H.; Passos, P.; Schleip, W.D. & Wüster, W. 2013. Best Practices: In the 21st Century, Taxonomic Decisions in Herpetology are Acceptable Only When Supported by a Body of Evidence and Published via Peer-Review. Herpetological Review 44 (1): 8-23
  • Kluge, Arnold G. 1993. Aspidites and the phylogeny of Pythonine snakes. Rec. Austral. Mus. (Supplement 19): 1-77 - get paper here
  • Knauf, S. 2000. Haltung und Nachzucht des Neu-Guinea-Wasserpythons (Liasis mackloti mackloti) (DUMÉRIL & BIBRON 1844). Elaphe 8 (4): 16-21
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  • McDiarmid, R.W.; Campbell, J.A. & Touré,T.A. 1999. Snake species of the world. Vol. 1. [type catalogue] Herpetologists’ League, 511 pp.
  • McDowell, S.B. 1975. A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part lI. Aniloidea and Pythoninae. Journal of Herpetology 9 (1): 1-79 - get paper here
  • Opferman, R. Regis 1994. The Savu Ialand python (Liasis mackloti savuensis). Reptiles 2 (1): 24-28
  • O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here
  • O’SHEA, Mark; Caitlin SANCHEZ, Andrew KATHRINER, Sven MECKE, Venancio LOPES CARVALHO, Agivedo VARELA RIBEIRO, Zito AFRANIO SOARES, Luis LEMOS DE ARAUJO and Hinrich KAISER 2015. Herpetological Diversity of Timor-Leste: Updates and a Review of Species Distributions. Asian Herpetological Research 6 (2): 73–131 - get paper here
  • O’SHEA, Mark; Caitlin SANCHEZ, Scott HEACOX, Andrew KATHRINER, Venancio LOPES CARVALHO, Agivedo VARELA RIBEIRO, Zito AFRANIO SOARES, Luis LEMOS DE ARAUJO and Hinrich KAISER 2012. First Update to Herpetofaunal Records from Timor-Leste. Asian Herpetological Research 3 (2): 114–126 - get paper here
  • PHILIPPEN, H.-D. & K.-D. SCHULZ 1994. Liasis mackloti savuensis BRONGERSMA 1956. Sauria 16 (1): 1-2 - get paper here
  • Radovanovic, Adam 2013. Captive management and reproduction of the Savu Island python Liasis mackloti savuensis (Brongersma, 1956). Herpetological Bulletin (123) - get paper here
  • Rawlings, L.H.; Barker, D.; Donnellan, S.C. 2004. Phylogenetic relationships of the Australo-Papuan Liasis pythons (Reptilia: Macrostomata), based on mitochondrial DNA. Australian Journal of Zoology 52 (2): 215-227 - get paper here
  • Reynolds, R. Graham; Matthew L. Niemiller, Liam J. Revell 2014. Toward a Tree-of-Life for the boas and pythons: Multilocus species-level phylogeny with unprecedented taxon sampling. Molecular Phylogenetics and Evolution 71: 201–213 [published online in 2013] - get paper here
  • Riel, C.A.P. van 1987. Breeding results: Liasis mackloti. Litteratura Serpentium 7 (3): 102 - get paper here
  • Schleip, Wulf D & O’Shea, M. 2010. Annotated checklist of the recent and extinct pythons (Serpentes, Pythonidae), with notes on nomenclature, taxonomy, and distribution. ZooKeys 66 (2010) : 29-79 - get paper here
  • Stimson, Andrew F. 1969. Liste der rezenten Amphibien und Reptilien: Boidae (Boinae + Bolyeriinae + Loxoceminae + Pythoninae). Das Tierreich 89 xi + 1-49
  • Swaak, H. 1988. Breeding with Liasis mackloti. Litteratura Serpentium 8 (6): 259-261 - get paper here
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  • Vidal, N., A.-S. Delmas, and S. B. Hedges 2007. The higher-level relationships of alethinophidian snakes inferred from seven nuclear and mitochondrial genes. Pp. 27-33 in R. W. Henderson and R. Powell (Eds.), Biology of the boas and pythons. Eagle Mountain Publishing, Eagle Mountain, Utah - get paper here
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  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
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