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Liolaemus porosus ABDALA, PAZ & SEMHAN, 2013

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Higher TaxaLiolaemidae, Iguania, Sauria, Squamata (lizards) 
Subspecies 
Common Names 
SynonymLiolaemus porosus ABDALA, PAZ & SEMHAN 2013 
DistributionArgentina (Salta), Chile (Atacama Region), above 3600 m elevation

Type locality: “20km al Este del puesto de Gendarmería, Argentina, ubicado en el límite internacional con Chile por el paso Socompa, departamento de Los Andes, provincia de Salta, Argentina. Coordenadas: 24°34’40.2’’ S y 68°11’58.4’’ W, 3707 msnm.” Map legend:
Type locality - Type locality.
 
Reproduction 
TypesHolotype: FML 2497, adult male 
CommentDiagnosis: Within the Liolaemus montanus series, L. porosus is the only species which shows postcloacal pores in males (1-8 x = 3.0 in five of the six males). It can be distin- guished from L. annectens; L. chlorostictus; L. dorbignyi; L. duellmani; L. fabiani; L. filiorum; L. forsteri; L. foxi; L. huayra; L. inti; L. jamesi ; L. melanogaster; L. nigriceps; L. orientalis; L. patriciaturrae, L. pachecoi ; L. pleopholis; L. polystictus; L. puritamensis; L. robustus, L. scrocchii; L. thomasi; L. vallecurensis; L. vulcanus, and L. williamsi, because all these species are larger (with a maximum snout- vent length in adults, between 77 and 105mm vs. 72.9mm in L. porosus). It differs from L. audituvelatus; L. erguetae; L. erroneus; L. ethe- ridgei; L. famatinae; L. fittkaui; L. insolitus; L. lopezi; L. manueli; L. ortizi; L. pantherinus; L. poconchilensis; L. reichei; L. rosenmanni; L. ruibali; L. schmidti; L. stolzmanni and L. torresi, because are smaller than L. porosus, (48-66mm maxSVL vs 72.9mm in L. porosus).
L. porosus is characterized by juxtaposed or subyuxtaposed, laminar, and without keel dorsal body scales; character states that diffe- rentiate it from L. disjunctus; L. etheridgei, L. fittkaui, L. huacahuasicus; L. montanus; L. orko; L. ortizi, L. polystictus, L. thomasi and L. tropidonotus which exhibit dorsal scales imbricate with evident keel. Dorsal scales also differentiate L. porosus from L. dorbignyi, L. famatinae, L. griseus; L. inti, L. huayra, L. jamesi, L. melanogaster, L. pleopholis, L. pulcherrimus, L signifer and L. williamsi which have imbricated or subimbricated and slightly keeled dorsal scales.
The number of scales around the body in L. porosus varies between 74-96 (x = 88.0), differing from several species of the group which have up to 73 scales, as in L. annectens, L. audituvelatus, L. chlorostictus, L. disjunctus, L. dorbignyi, L. etheridgei, L. fabiani, L. fama- tinae, L. fittkaui, L. griseus, L. huacahuasicus, L. huayra, L. insolitus, L. islugensis; L. jame- si, L. lopezi, L. melanogaster, L. montanus, L. orientalis, L. orko, L. ortizi, L. pachecoi, L. poconchilensis, L. polystictus, L. puritamensis, L. robustus, L. scrocchii, L. stozlmanni, L. tho- masi, L. torresi, L. tropidonotus, L. vulcanus, and L. williamsi.
The number of dorsal scales between occi- put and hind limbs in L. porosus varies between 89 and 104 (x = 95.6), and is higher than L. annectens, L. audituvelatus, L. chlorostictus, L. dorbignyi, L. etheridgei, L. fittkaui, L. griseus, L. huacahuasicus, L. huayra, L. inti, L. islugen- sis, L. jamesi, L. melanogaster, L. montanus, L. orientalis, L. orko, L. pachecoi, L. pocon- chiliensis, L. polystictus, L. puritamensis, L. robustus, L. scrocchii, L. thomasi, L. vulcanus, and L. williamsi, which this scales are not higher than 88.
The number of ventral scales between mental and ventral edge of the cloaca in L. porosus is higher than in L. annectens, L. chlorostictus, L. etheridgei, L. fabiani, L. famatinae, L. fittkahui, L. forsteri, L. griseus, L. huacahuasicus, L. huayra, L. islugensis, L. jamesi, L. melanogaster, L. montanus, L. orientalis, L. pachecoi, L. polystictus, L. pul- cherrimus, L. puritamensis, L. robustus, L. ruibali, L. signifer, L. thomasi, and L. williamsi the (92-109; x = 99.8 vs up to 91, respectively).
In Liolaemus porosus the number of sca- les in neck (from posterior edge of auricular meatus to shoulder, along the longitudinal fold) varies between 47-51 (x = 49.0), being higher than L. annectens, L. audituvelatus, L. dorbignyi, L. eleodori; L. fabiani, L. famatinae, L. fittkahui, L. forsteri, L. foxi, L. halonastes, L. huacahuasicus, L. islugensis, L. jamesi, L. melanogaster, L. montanus, L. orientalis, L. orko, L. pachecoi, L. puritamensis, L. robertoi; L. robustus, L. ruibali, L. scrocchii, L. schmidti, L. signifer, L. thomasi, L. torresi, and L. vul- canus, which have up to 45 neck’s scales.
The number of gular scales is higer in Liolaemus porosus than in L. annectens, L. chlorostictus, L. erguetae, L. etheridgei, L. fabiani, L. famatinae, L. fittkaui, L. griseus, L. huacahuasicus, L. huayra, L. jamesi, L. mela- nogaster, L. montanus, L. orientalis, L. orko, L. pachecoi, L. pleopholis, L. polystictus, L. pulcherrimus, L. robertoi, L. robustus, L. signi- fer, L. thomasi, and L. torresi (41-52; x = 46.1; vs up to 40, respectively).
Number of precloacal pores in males in Liolaemus porosus (5-7; x=5.8) is higher than in the L. halonastes, L. manueli, and L. pocon- chiliensis (x=<4.8)
The presence of precloacal pores in fema- les (4-5; x = 4.4) distinguish the new taxon from L. audituvalatus, L. filiorum, L. fittkaui, L. forsteri, L. hajeki, L. halonastes, L. jamesi, L. lopezi, L. melanogaster, L. molinai; L. ortizi, L. poconchiliensis, L. polystictus, L. purita- mensis, L. robertoi, L. ruibali, L. torresi, and L. vallecurensis which lack precloacal pores. Also, differ from females with up to four (x <3.0) precloacal pores: L. andinus (x = 2.0), L. annectens (x = 0.5), L. eleodori (x = 2.7), Liolaemus gracielae (x = 0.3), L. multicolor (x = 0.1), L. orientalis (x = 1.8), and L. poecilo- chromus (x = 0.3).
The number of supernumerary pores of males of Liolaemus porosus (1-12; x = 5.5) differ from all species of the L. montanus group which lack supernumerary pores, with the exception of L. cazianae; L. famatinae, L. forsteri, L. griseus, L. huayra, L. melanogas- ter, L. orientalis, L. orko, L. pulcherrimus, L. signifer and L. thomasi, which exhibit super- numerary pores, but they have a lower number of pores (maximum datum registered from L. griseus x = 2.3). 
Etymologynamed after Latin (Porosus = porous) referring to the large number of pre-cloacal and post-cloacal pores. 
References
  • Abdala, Cristian Simón; Marcos Maximiliano Paz & Romina Valeria Semhan 2013. Nuevo Liolaemus (Iguania: Liolaemidae) con novedoso carácter morfológico, de la frontera entre Argentina y Chile. Rev. Biol. Trop. 61 (4): 1563-1584 - get paper here
  • Demangel, Diego; Cristian Sepúlveda, Manuel Jara, Daniel Pincheira-Donoso<br />and Herman Núñez 2015. Liolaemus omorfi, A NEW LIZARD SPECIES FROM THE ANDES OF NORTHERN CHILE (SAURIA, LIOLAEMIDAE). Boletín del Museo Nacional de Historia Natural, Chile, 64: 139-155 - get paper here
  • Kwet, Axel 2014. Liste der im Jahr 2013 neu beschriebenen Reptilien. Terraria Elaphe 2014 (3): 56-67 - get paper here
 
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