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Liolaemus tromen ABDALA, SEMHAN, MORENO AZOCAR, BONINO, PAZ & CRUZ, 2012

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Higher TaxaLiolaemidae, Iguania, Sauria, Squamata (lizards) 
Subspecies 
Common Names 
SynonymLiolaemus tromen ABDALA, SEMHAN, MORENO AZOCAR, BONINO, PAZ & CRUZ 2012
Liolaemus hermannunezi — ABDALA & QUINTEROS 2007
Liolaemus hermannunezi — TULLI et al. 2009 
DistributionArgentina (Neuquén)

Type locality: Km 140 of provincial route 21.7 km North of Huecú, El Cholar departament, Neuquén Province, Argentina  
Reproduction 
TypesHolotype: FML 17735. C. Abdala, C. Robles and R. Juárez Cols. 21/I/2006. Paratypes.— FML 17731 – 734 and FML 17736: Same locality and date of the holotype. 
CommentSimilar species: confused initially with Liolaemus hermannunezi, a mountain Chilean species.

Diagnosis. Liolaemus tromen belongs to the L. melanops group. Within this group, the new species differenciates from L. boulengeri, L. donosobarrosi, L. goetschi, L. hermannunezi, L. inacayali, L. josei, L. loboi, L. martorii, L. melanops, L. puelche, L. rothi, L. sagei, L. senguer, L. tehuelche and L. telsen, because it shows a short antehu- meral band that is not longer than the superior line of forelimbs, absent in the above-mentioned species. It also dif- ferenciates from the species belonging to the L. telsen group (except for L. rothi and L. sagei), because L. tromen has a larger snout-vent length (SVL) of 82.5 mm versus the SVL in the species of L. telsen group (62.3 – 77.1 mm).
It is different from Liolaemus mapuche by having four scales contacting mental scale (four to six in L. mapuche), by having a more expanded melanic ventral surface and by the absence of light blue scales on mid body and tail. It is different from L. cuyanus, because it is smaller (max SVL 82.5 mm, whereas L. cuyanus is 102 mm), four scales in contact with mental scale (six in L. cuyanus) and by a broader ventral melanism.
The new species segregates from the Liolaemus fitzingerii group (L. fitzingerii, L. canqueli, L. casamiquelai, L. chehuachekenk, L. fitzingerii, L. morenoi and L. xanthoviridis), by its strong sexual dichromatism, absent in all species except for L. melanops. Moreover, it is also remarkably shorter in SVL (maximum SVL 82.5 mm in L. tromen vs. 88.4–110 mm in the other species), has a short antehumeral band, never wide or expanded and never going further than the superior line of the forelimbs. Liolaemus tromen is different from L. canqueli, L. casamique- lai, L. fitzingerii, and L. xanthoviridis because it shows evident pre and postscapular spots. Comparing it to L. melanops, L. tromen is differnt by lacking cephalic melanism, the presence of the mentioned pre and postscapular spots and males are never green in L. tromen.

Group: member of the Liolaemus rothi complex. 
EtymologyThe species is named after the Tromen Volcano, Neuquén Province, Argentina. Tromen means “totora” or “cloudy or shady place” in the mapuche language. 
References
  • ABDALA C. S., R. V. SEMHAN, D. L. MORENO AZOCAR, M. BONINO, M. M. PAZ & F. CRUZ 2012. Taxonomic study and morphology based phylogeny of the patagonic clade Liolaemus melanops group (Iguania: Liolaemidae), with the description of three new taxa. Zootaxa 3163: 1–32 - get paper here
  • AVILA, LUCIANO JAVIER; LORENA ELIZABETH MARTINEZ & MARIANA MORANDO 2013. Checklist of lizards and amphisbaenians of Argentina: an update. Zootaxa 3616 (3): 201–238
  • Kwet, Axel 2013. Liste der im Jahr 2012 neu beschriebenen Reptilien. Terraria-Elaphe 2013 (3): 52-67 - get paper here
  • LUCIANO JAVIER AVILA, MELISA OLAVE, CRISTIAN HERNAN FULVIO PEREZ, DANIEL ROBERTO PEREZ, MARIANA MORANDO 2013. Molecular phylogenetic relationships of the Liolaemus rothi complex and a new species of lizard from Auca Mahuida Volcano (Squamata: Liolaemini). Zootaxa 3608 (4): 221–238 - get paper here
  • Olave, Melisa; Luciano J. Avila, Jack W. Sites Jr, Mariana Morando 2016. Hidden diversity within the lizard genus Liolaemus: genetic vs morphological divergence in the L. rothi complex (SQUAMATA: LIOLAEMINAE). Molecular Phylogenetics and Evolution - get paper here
 
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