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Liopholis multiscutata (MITCHELL & BEHRNDT, 1949)

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Higher TaxaScincidae, Egerniinae, Scincoidea, Sauria, Squamata (lizards) 
SubspeciesLiopholis multiscutata multiscutata (MITCHELL & BEHRNDT 1949)
Liopholis multiscutata bos (STORR 1960) 
Common NamesSouthern Sand-skink, Heath Skink, Bull Skink 
SynonymEgernia whitii multiscutata MITCHELL & BEHRNDT 1949: 176
Egernia bos STORR 1960
Egernia bos — FORD 1963
Egernia multiscutata — COGGER 1983: 158
Egernia multiscutata — COGGER 2000: 468
Egernia multiscutata — COUPER et al. 2006: 380
Liopholis multiscutata — GARDNER et al. 2008
Liopholis multiscutata — WILSON & SWAN 2013: 330

Liopholis multiscutata bos (STORR 1960)
Egernia multiscutata bos STORR 1960
Egernia multiscutata bos — STORR et al. 1999 
DistributionAustralia (South Australia: Kangaroo Island and Eyre and Yorke Peninsulas to coastal regions of S Western Australia, W Victoria, many offshore islands)

multiscutata: Greenly Island, South Australia.
bos: remainder of range.

Type locality: Greenly Is., S. A.  
ReproductionViviparous. 
TypesHolotype: SAMA R2636
Holotype: WAM R10751, from Cheyne Beach, 32 mi E of Albany, W. A. [bos] 
CommentSynonymy after COGGER 1983.

Subspecies: DONNELLAN et al. (2002) rejected the recognition of subspecies based on genetic and morphological evidence.

Habitat: Obligate burrowing.

This species exhibit colour pattern polymorphism (Chapple et al. 2008). 
References
  • Brown, Geoff W.; Peter Robertson and Ben G. Fanson 2018. Digging in: a review of the ecology and management of a threatened reptile with a small disjunct distribution – the heath skink, Liopholis multiscutata, in Victoria, south-eastern Australia. Pacific Conservation Biology - get paper here
  • Chapple, D. G., Hutchinson, M.N., Maryan, B., Plivelich M., Moore, J.A. and Keogh J.S. 2008. Evolution and maintenance of colour pattern polymorphism in Liopholis (Squamata : Scincidae). Australian Journal of Zoology 56: 103–115 - get paper here
  • CHAPPLE, DAVID G. and J. SCOTT KEOGH 2004. Parallel adaptive radiations in arid and temperate Australia: molecular phylogeography and systematics of the Egernia whitii (Lacertilia: Scincidae) species group. Biological Journal of the Linnean Society 83: 157–173 - get paper here
  • Chapple, David G.; J. Scott Keogh and Mark N. Hutchinson 2004. Molecular phylogeography and systematics of the arid-zone members of the Egernia whitii (Lacertilia: Scincidae) species group. Molecular Phylogenetics and Evolution 33 (3): 549-561 - get paper here
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp.
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Couper, P., Covacevich, J., Amey, A. & Baker, A. 2006. The genera of skinks (Family Scincidae) of Australia and its island territories: diversity, distribution and identification. in: Merrick, J.R., Archer, M., Hickey, G.M. & Lee, M.S.Y. (eds.). Evolution and Zoogeography of Australasian Vertebrates. Australian Scientific Publishing, Sydney, pp. 367-384
  • Donnellan,S.C.; Hutchinson M N., Dempsey, P. & Osborne, W.S. 2002. Systematics of the Egernia whitii species group (Lacertilia: Scincidae) in south-eastern Australia. Australian Journal of Zoology 50: 439-459 - get paper here
  • Ford, J. 1963. The Reptilian Fauna of the Islands between Dongara and Lancelin, Western Australia. Western Australian Naturalist 8 (6): 135-142
  • Gardner, M. G., Hugall, A. F., Donnellan, S. C., Hutchinson, M. N., and Foster, R. 2008. Molecular systematics of social skinks: phylogeny and taxonomy of the Egernia group (Reptilia: Scincidae). Zoological Journal of the Linnean Society, 154 (4): 781-794 - get paper here
  • Mitchell, F. J. & Behrndt, A. C. 1949. Fauna and flora of the Greenly Islands. Part 1. Introductory narrative and vertebrate fauna. (Reptiles: 175-178). Rec. South Austral. Mus. 9: 167-179 - get paper here
  • Storr G M 1978. The genus Egernia (Lacertilia, Scincidae) in Western Australia. Rec. West. Aust. Mus. 6 (2): 147-187 - get paper here
  • Storr, G. M. 1960. Egernia bos, a new skink from the south coast of Western Australia. Western Australian Naturalist 7: 99-103
  • Storr, G. M. 1968. Revision of the Egernia whitei species-group (Lacertilia: Scincidae). J. R. Soc. West. Aust. 51: 51-62
  • Storr, G. M., Smith, L. A. & Johnstone, R. E. 1981. Lizards of Western Australia. I. Skinks. Perth: University of Western Australia Press and Western Australian Museum, 200 pp.
  • Storr, G. M.; L. A. Smith, and R. E. Johnstone 1999. Lizards of Western Australia. I. Skinks. Revised Edition. Western Australian Museum
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
 
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