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Lophognathus gilberti GRAY, 1842

IUCN Red List - Lophognathus gilberti - Least Concern, LC

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Higher TaxaAgamidae (Amphibolurinae), Sauria, Iguania, Squamata (lizards) 
Subspecies 
Common NamesE: Gilbert's Lashtail, Gilbert's Dragon
G: Gilberts Agame 
SynonymLophognathus gilberti GRAY 1842: 53
Redtenbacheria fasciata STEINDACHNER 1867: 31
Grammatophora temporalis GÜNTHER 1867: 52 (part.)
Lophognathus gilberti — BOULENGER 1883: 225
Physignathus gilberti — BOULENGER 1885: 396
Physignathus incognitus AHL 1926
Gemmatophora gilberti — STORR 1983
Amphibolurus gilberti — EHMANN 1992
Lophognathus gilberti — COGGER 2000: 340
Amphibolurus gilberti — GREER 2006
Amphibolurus gilberti — WILSON & SWAN 2010: 350
Lophognathus gilberti — MELVILLE et al. 2011
Amphibolurus gilberti — MAYER 2014
Lophognathus gilberti — MELVILLE et al. 2018 
DistributionAustralia (Northern Territory, W/C Queensland, N/NW Western Australia)

Type locality: Port Essington, N. T.  
Reproductionoviparous 
TypesHolotype: BMNH 1946.8.28.69
Paralectotype: BMNH 1946.8.12.73 from Nickol Bay, W. A. [G. temporalis]
Holotype: NMW, but apparently not found fide Melville et al. 2018 [Redtenbacheria fasciata]
Holotype: ZMB 30086, from Australia [incognitus] 
DiagnosisDiagnosis (genus): A genus consisting of large agamid lizards in the subfamily Amphibolurinae, with exposed tympanum, gular scales smooth to weakly keeled, ventral scales weakly to strongly keeled. Stoutly built with moderately long legs and tail. Broad white stripe on the upper and lower lips, extending along the full extent of the jaw, pale stripe from behind the eye to the top of the ear, which is cream, white, grey or yellow in life. This pale stripe is well defined ventrally and dorsally by a row of darkly pigmented scales (fig. 6). Heterogenous scales on the back both at the midline and doroslaterally, associated with a weak to prominent row of enlarged, strongly keeled scales. Colour patterns of grey, brown, rust-brown and black. Well- developed nuchal crest continuous with the enlarged row of vertebral scales. Broad pale dorsolateral stripes, which may extend from top of ear or back of head to hips. Dorsolateral stripes are not continuous with the pale lip stripes. On the back, dorsolateral stripes may be intersected by wedges of brown or grey. Preanal pores 3–6; femoral pores 2–8 (Melville et al. 2018: 53)

Diagnosis (gilberti): As for genus. Lophognathus gilberti is distinguished from Lophognathus horneri sp. nov. by lacking a distinct white spot on the tympanum (fig. 7) that is surrounded by or adjacent to black pigmentation (Melville et al. 2018: 54). 
CommentType species: Lophognathus gilberti GRAY 1842 is the type species of the genus Lophognathus GRAY 1842.

Synonymy partly after COGGER 1983. Physignathus gilberti centralis LOVERIDGE 1932 has been removed from the synonymy of L. gilberti by WELLS & WELLINGTON 1984. 
EtymologyNamed after the British naturalist John Gilbert, who collected some of the specimen on which their first description is based. 
References
  • Ahl, E. 1926. Neue Eidechsen und Amphibien. Zool. Anz. 67: 186-192
  • Beolens, ; Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, G. A. 1883. Remarks on the lizards of the genus Lophognathus. Ann. Mag. Nat. Hist. (5) 12: 225-226 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp.
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Ellis, M.; Higgins, D. 1993. The Occureence of Gillbert's Dragon Lophognathus gilberti in the Paroo Drainage Basin, Western New South Wales. In Lunney & Ayers: Herpetology In Australia 1993: 35-36
  • Gray, J. E. 1842. Description of some hitherto unrecorded species of Australian reptiles and batrachians. Zoological Miscellany 2: 51—57 (London: Treuttel, Würtz & Co) - get paper here
  • Günther, A. 1867. Additions to the knowledge of Australian reptiles and fishes. Ann. Mag. nat. Hist. (3) 20: 45-57 - get paper here
  • Houston, T. F. 1978. Dragon Lizards and Goannas of South Australia. Adelaide: South Australian Museum 84 pp.
  • Loveridge, A. 1933. New agamid lizards of the genera Amphibolurus and Physignathus from Australia. Proc. New England zool. Club 13: 69-72
  • Mayer, M. 2014. Von Schlangen, Kröten und Krokodilen im tropischen „Top End“ Australiens. Ein Reise- und Studienbericht. Reptilia (Münster) 19 (110): 76-85
  • Melville, J.; E.G. Ritchie; S.N.J. Chapple, R.E. Glor, J.A. Schulte II 2011. Evolutionary origins and diversification of dragon lizards in Australia’s tropical savannas. Molecular Phylogenetics and Evolution 58: 257–270 - get paper here
  • MELVILLE, JANE; EUAN G. RITCHIE, STEPHANIE N.J. CHAPPLE, RICHARD E. GLOR AND JAMES A. SCHULTE 2018. Diversity in Australia’s tropical savannas: An integrative taxonomic revision of agamid lizards from the genera Amphibolurus and Lophognathus (Lacertilia: Agamidae). Memoirs of Museum Victoria 77: 41–61 - get paper here
  • Steindachner, F. 1867. In: Reise der Österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858,1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair (Zoologie), Vol. 1, part 3 (Reptilien p.1-98). K. Gerold's Sohn/Kaiserlich-Königl. Hof- und Staatsdruckerei, Wien [1869 on title page] - get paper here
  • Storr, G. M. 1974. Agamid lizards of the genera Caimanops, Physignathus and Diporiphora in Western Australia and Northern Territory. Rec. West. Aust. Mus. 3: 121-146 - get paper here
  • Thompson G G & S A Thompson 2001. Behavioural and spatial ecology of Gilbert's dragon Lophognathus gilberti (Agamidae: Reptilia). J. Royal Soc. Western Australia 84: 153-158
  • Werning, H. 2002. Wasseragamen und Segelechsen. Natur und Tier Verlag (Münster), 127 pp. [review in Sauria 26 (4): 17] - get paper here
  • Werning, H. 2004. Bibliographie der Gattungen Physignathus, Lophognathus und Hydrosaurus. Iguana Rundschreiben 17 (2): 18-31
  • Werning,H. 1995. Wasseragamen [water dragons]. Terrarien Bibliothek, Heselhaus & Schmidt Verlag, Münster [ISBN 3-9801853-9-7]
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
 
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