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Lygosoma schneideri WERNER, 1900

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Higher TaxaScincidae, Lygosominae, Scincoidea, Sauria, Squamata (lizards) 
Subspecies 
Common NamesE: Sumatran Supple Skink
G: Sumatra Laubskink 
SynonymLygosoma (Riopa) schneideri WERNER 1900: 487
Lygosoma bampfyldii — BOULENGER 1912: 93 (in part)
Lygosoma bampfyldei — DE ROOIJ 1915: 263 (in part)
Lygosoma bampfyldii — SMITH 1930: 36 (in part)
Riopa bampfyldei — SMITH 1937: 228
Mochlus bampfyledei — MITTLEMAN 1952: 22.
Lygosoma bampfyldei — GREER 1977: 915
Lygosoma bampfyldei — WELCH et al. 1990: 83
Riopa bampfyldii — DENZER & MANTHEY 1991: 317
Riopa bampfyldei — MANTHEY & GROSSMANN 1997: 275
Riopa bampfyldei — CHAN-ARD et al. 1999: 27
Riopa bampfyldei — MALKMUS et al. 2002: 281
Lygosoma bampfyldei — DAS & YAAKOB 2007
Lygosoma bampfyldei — DAS 2010 238
Lygosoma bampfyldei — GRISMER 2008
Lygosoma bampfyldei — GRISMER 2011: 613 
DistributionIndonesia (Sumatra)

Type locality: “Indragiri, Djapura”, Sumatra, Indonesia  
Reproduction 
TypesHolotype: NMBA 4743, Adult male, accessioned into the Naturhistorisches Museum Basel by G. Schneider on 1 January 1900. 
DiagnosisDiagnosis. Lygosoma schneideri can be differentiated from all other Lygosoma by having the combination of a relatively large (SVL = 129 mm) robust body (PEC/SVL = 0.15); seven supralabials and infralabials; midline contact of the supranasals; prefrontals not in contact; frontoparietal contacting three supraoculars; large postinterparietal present; eight superciliaries; two postsuboculars, the first being small; one or two primary and three secondary and tertiary temporals; eight nuchal scales; a deep postnasal groove extending from the nasal scale to below the anterior portion of the eye and lying below the anterior loreals and lower preocular and above the second and third supralabials; scaly lower eyelid, no window; 45 midbody scale rows; 95 paravertebral scale rows; 98 ventral scale rows; 34 caudal scale rows at the tenth subcaudal; eight large precloacal scales; smooth to weakly keeled subdigital finger lamellae, 10 lamellae on third finger; keeled subdigital toe lamellae, 16 lamellae on fourth toe; low, round, small, palmar scales numbering seven across the base of the palm; head pattern consisting of a dark, continuous frontal and occipital band separated by a yellowish band; dark occipital band not confluent with lighter color of dorsum and tail. These characters are scored across all species in the L. bampfyldei group in Table 2 (Grismer et al. 2018). 
CommentSynonymy: synonymized with L. bampfyldei by KRAMER 1979 but resurrected from synonymy by Grismer et al. 2018.

Distribution: Das (2010) reports this species from “north-western Sumatra (Indrajiri, Riau Province)” however the type locality listed by Werner (1910) “Indragiri, Djapura” is in southeastern Sumatra (Fig. 1 in Grismer et al. 2018). 
Etymology 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, G.A. 1912. A vertebrate fauna of the Malay Peninsula from the Isthmus of Kra to Singapore incl. the adjacent islands. Reptilia and Amphibia. London (Taylor & Francis), xiii + 298 S. - get paper here
  • Chan-ard,T.; Grossmann,W.; Gumprecht,A. & Schulz,K. D. 1999. Amphibians and reptiles of peninsular Malaysia and Thailand - an illustrated checklist [bilingual English and German]. Bushmaster Publications, Würselen, Gemany, 240 pp. [book review in Russ. J Herp. 7: 87] - get paper here
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • Das, I. & Yaakob, N. 2007. Status of knowledge of the Malaysian herpetofauna. In Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. in: L. S. L. Chua, L. G. Kirton & L. G. Saw (eds.), Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. Forest Research Institute Malaysia, Kepong, pp. 31-81
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp.
  • Denzer, W. & MANTHEY, U. 1991. A nominal checklist of the lizards inhabiting Peninsular Malaysia and Singapore. Raffles Bull. Zool., 39(2): 309-322 - get paper here
  • GEISSLER, PETER; TIMO HARTMANN & THY NEANG 2012. A new species of the genus Lygosoma Hardwicke & Gray, 1827 (Squamata: Scincidae) from northeastern Cambodia, with an updated identification key to the genus Lygosoma in mainland Southeast Asia. Zootaxa 3190: 56–68
  • Greer, A.E. 1977. The systematics and evolutionary relationships of the scincid lizard genus Lygosoma. Journal of Natural History 11: 515-540.
  • Grismer, L. Lee 2008. A revised and updated checklist of the lizards of Peninsular Malaysia. Zootaxa 1860: 28–34 - get paper here
  • Grismer, L. Lee; Chan K. Onn, Jesse L. Grismer, Perry L. Wood, Jr., and A. Norhayati 2010. A CHECKLIST OF THE HERPETOFAUNA OF THE BANJARAN BINTANG, PENINSULAR MALAYSIA. Russ. J. Herpetol. 17 (2): 147-160 - get paper here
  • GRISMER, L. LEE; EVAN S. H. QUAH, ZAHARIL DUZULKAFLY & PAUL YAMBUN 2018. On the taxonomy of Lygosoma bampfyldei Bartlett, 1895 (Squamata: Scincidae) with descriptions of new species from Borneo and Peninsular Malaysia and the resurrection of Lygosoma schneideri Werner, 1900. Zootaxa 4438 (3): 528–550 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Kramer, Eugen 1979. Typenkatalog der Echsen im Naturhistorischen Museum Basel (BM), Stand 1978. [type catalogue] Revue Suisse de Zoologie 86 (1): 159-166 - get paper here
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Mittleman, M.B. 1952. A generic synopsis of the Lizards of the Subfamily Lygosominae. Smithson. misc. Collns, 117: 1-35. - get paper here
  • Smith,M.A. 1930. The Reptilia and Amphibia of the Malay Peninsula fromt he Isthmus of Kra to Singapore, including the adjacent Islands. A Supplement to G.A. Boulenger’s Reptilia and Batrachia 1912. Bull. Raffles Mus. No. 3: 1-149
  • Smith,M.A. 1937. A review of the genus Lygosoma (Scincidae: Reptilia) and its allies. Records of the Indian Museum 39 (3): 213-234
  • Welch, K.R.G., Cooke, P.S., & Wright, A.S. 1990. Lizards of the Orient: a checklist. Robert E. Krieger Pub. Co., Malabar, FL 162 pp.
  • Werner, F. 1900. Reptilien und Batrachier aus Sumatra, gesammelt Herrn. Gustav Schneider jr., im Jahre 1897-1898. Zool. Jahrb. (syst) 13: 479-508 - get paper here
 
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