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Microlophus arenarius (TSCHUDI, 1845)

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Higher TaxaTropiduridae, Iguania, Sauria, Squamata (lizards)
Common Names 
SynonymSteironotus (Eulophus) arenarius TSCHUDI 1845: 154
Tropidurus stolzmanni STEINDACHNER 1891: 376
Tropidurus tschudii ROUX 1907
Tropidurus occipitalis stolzmanni — MERTENS 1956
Ophryoessoides arenarius — ETHERIDGE 1966
Tropidurus arenarius — FRITTS 1974
Tropidurus stolzmanni — DIXON & WRIGHT 1975: 34
Microlophus stolzmanni — FROST 1992
Tropidurus occipitalis stolzmanni — TIEDEMANN et al. 1994
Tropidurus arenarius — LEHR 2002: 203
Stenocercus arenarius — TORRES-CARVAJAL 2004
Microlophus stolzmanni — TORRES-CARVAJAL 2004
Tropidurus arenarius — TORRES-CARVAJAL 2005 (pers. comm.)
Microlophus arenarius — CARVALHO 2021 

Type locality: Huacho, north of Lima, Peru.

stolzmanni: N Peru; Type locality: “bei Chota”  
Reproductionoviparous; <br />stolzmanni: Females from 65 to 76 mm in snout-vent length contained 3 to 4 oviducal eggs. All oviducal eggs examined were shelled and apparently ready for deposition. One female 73 mm in snout-vent length contained 16 unyolked ovarian eggs. These lizards were taken between 23-28 November 1969 [DIXON & WRIGHT 1975]. 
TypesLectotype: MHNN 2275, designated by Carvalho 2021
Syntypes: NMW 18908: 1, 2 [stolzmanni] 
DiagnosisDiagnosis. Tropidurus stolzmanni is a member of the occipitalis group and differs from the other members of the group by lacking middorsal spots and by having chevron-shaped lines in the gular region, more prominent in males; differing also from occipitalis in lacking an occipital spot. Adult males of stolzmanni differ from males of both koepckeorum and occipitalis in having a broad, black-bordered, white stripe from the eye to near midbody [DIXON & WRIGHT 1975]. 
CommentSynonymy: fide Carvalho 2021.

Habitat. The majority of specimens were found on or near small boulders, trees, and shrubs. Several were found under debris associated with abandoned human dwellings. Most were found in habitats with some kind of vegetation, such as mesquite, acacia, Capparis scabrida, dense beach shrubs or semiarid woodlands of the foothill slopes. This species is primarily scansorial; most specimens were taken one to three dm above the ground on trees. However, some were taken on the ground in the vicinity of shrubs where they appeared to be foraging for food [DIXON & WRIGHT 1975 on M. stolzmanni].

NCBI taxonID: 179259 [stolzmanni] 
  • Bauer, A.M.; Günther,R. & Klipfel,M. 1995. The herpetological contributions of Wilhelm C.H. Peters (1815-1883). SSAR Facsimile Reprints in Herpetology, 714 pp.
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Boulenger, George A. 1901. Further descriptions of new reptiles collected by Mr. P. O. Simons in Peru and Bolivia. Ann. Mag. nat. Hist. (7) 7 (42): 546-549 - get paper here
  • Carvalho, André L.G. 2021. Resolving the Obscure Identity of Steironotus arenarius Tschudi, 1845 and Tropidurus tschudii Roux, 1907 (Squamata: Tropiduridae). South American Journal of Herpetology 19(1), 40-53 - get paper here
  • Cuadrado SS, Loor YA, Narváez AE 2020. Herpetofauna of Engabao, Playas Canton, Ecuador, with notes on the occurrence of Ceratoprhys stolzmanni (Steindachner, 1882). Check List 16(3): 665-674 - get paper here
  • Dixon, J.R. & WRIGHT,J.W. 1975. A review of the lizards of the iguanid genus Tropidurus in Peru. Contributions in Science (Los Angeles) (271): 1-39 - get paper here
  • Etheridge, Richard E. 1966. The systematic relationships of West Indian and South American lizards referred to the iguanid genus Leiocephalus. Copeia 1966 (1): 79-91 - get paper here
  • Fritts, Thomas H. 1974. A multivariate evolutionary analysis of the Andean Iguanid lizards of the genus Stenocercus. San Diego Society of Natural History Memoir 7: 1-89
  • Goldberg, S.R. & Bursey, C.R. 2009. Helminths from seven species of Microlophus (Squamata: Tropiduridae). Salamandra 45 (2): 125-128 - get paper here
  • HARVEY, Michael B. & RONALD L. GUTBERLET JR 2000. A phylogenetic analysis of the tropidurine lizards (Squamata: Tropiduridae), including new characters of squamation and epidermal microstructure. Zoological Journal of the Linnean Society 128: 189–233. - get paper here
  • Koch, C. 2010. Geheimnisvolles Peru. Reptilia (Münster) 15 (82): 36-45 - get paper here
  • Lehr, E. 2002. Amphibien und Reptilien in Peru. Natur und Tier-Verlag (Münster), 208 pp. - get paper here
  • Müller, L. 1924. Ueber neue oder seltene Mittel- und Südamerikanische Amphibien und Reptilien. Mitt. Zool. Mus. Berlin 11 (1): 75-93 [1923]
  • Peters, Wilhem Carl Hartwig. 1871. Über einige Arten der herpetologischen Sammlung des Berliner zoologischen Museums. Monatsber. Preuss. Akad. Wiss. Berlin 1871: 644-652 - get paper here
  • Schlüter, U. 2000. Wüstenkielschwanzleguane aus Peru. Draco 1 (4): 44-49
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
  • Torres-Carvajal, O. 2004. Herpetofauna of Isla de La Plata, Ecuador. Herpetological Review 35 (1): 85 - get paper here
  • Torres-Carvajal, O. 2004. The abdominal skeleton of Tropidurid lizards (Squamata: Tropiduridae). Herpetologica 60 (1): 75-83 - get paper here
  • Toyama, Ken Sebastian 2016. Evidence of adaptive evolution in the cranial morphology of Tropidurid lizards from coastal Peru. Herpetology Notes 9: 47-53 - get paper here
  • Tschudi,J. J. von 1845. Reptilium conspectum quae in republica Peruana reperiuntur er pleraque observata vel collecta sunt in itenere. Archiv für Naturgeschichte 11 (1): 150-170 [reprint 1968, SSAR; sometimes cited as being published in 1846]. - get paper here
  • Watkins, G. W. 1996. Proximate causes of sexual size dimorphism in the iguanian lizard Microlophus occipitalis. Ecology 77: 1473-1482 - get paper here
  • Watkins, Graham G. 1998. Function of a secondary sexual ornament: the crest in the South American Iguanian lizard Microlophus occipitalis (Peters, Tropiduridae). Herpetologica 54 (2): 161-169 - get paper here
  • Werner, FRANZ 1901. Reptilien und Batrachier aus Peru und Bolivien. Abhandl. Ber. Zool. Anthrop. Mus. Dresden, 9 (2): 1-14 [1900] - get paper here
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