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- Benavides, Edgar; Rebecca Baum, Heidi M. Snell, Howard L. Snell, and Jack W. Sites, Jr. 2009. Island Biogeography of Galápagos Lava Lizards (Tropiduridae: Microlophus): Species Diversity and Colonization of the Archipelago. Evolution 63 (6): 1606-1626, doi: 10.1111/j.1558-5646.2009.00617.x - get paper here
- HervÍas-Parejo, Sandra; Ruben Heleno, Beatriz Rumeu, Beatriz Guzmán, Pablo Vargas, Jens M Olesen, Anna Traveset, Carlos Vera, Edgar Benavides, Manuel Nogales, 2019. Small size does not restrain frugivory and seed dispersal across the evolutionary radiation of Galápagos lava lizards. Current Zoology, Volume 65, Issue 4, August , Pages 353–361, - get paper here
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