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Pachydactylus formosus SMITH, 1849

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Southern Rough Gecko, Karoo Gecko, Smith's Thick-toed Gecko 
SynonymPachydactylus formosus SMITH 1849
Pachydactylus mento-marginatus SMITH 1849 (fide LOVERIDGE 1947)
Pachydactylus formosus — BOULENGER 1885: 203
Pachydactylus mentomarginatus — BOULENGER 1885: 207
Pachydactylus obscurus THOMINOT 1889: 21 (fide LOVERIDGE 1947)
Pachydactylus capensis formosus — HEWITT 1927: 400
Pachydactylus capensis formosus — LOVERIDGE 1947
Pachydactylus capensis formosus — WERMUTH 1965: 117
Pachydactylus rugosus formosus — JOGER 1985: 480
Pachydactylus rugosus formosus — BRANCH 1998: 259
Pachydactylus rugosus formosus — RÖSLER 2000: 99
Pachydactylus formosus — LAMB & BAUER 2000
Pachydactylus formosus — BATES et al. 2014: 129
Pachydactylus formosus — SCHLEICHER 2020 
DistributionRepublic of South Africa (S Little Namaqualand, Cape Province), Namibia, Botswana

Type locality: Interior of South Africa near the Tropic of Capricorn.  
Reproductionoviparous 
TypesHolotype: BMNH 1946.8.30.71 
Diagnosis 
CommentP. formosus is listed by KLUGE (1993) as a valid species with question mark. LAMB & BAUER (2000) revalidated formosus to full species status. Distribution not yet corrected for elevation of P. rugosus formosus to species status. 
References
  • Auerbach, R.D. 1987. The Amphibians and Reptiles of Botswana. Mokwepa Consultants, Botswana, 295 pp.
  • Barts, M. 2012. Die Dickfingergeckos des südlichen Afrikas. Teil XIII: Karoo Dickfingergecko Pachydactylus formosus SMITH, 1849. Sauria 34 (1): 37-42 - get paper here
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Branch, W. R. 1998. Field Guide to the Snakes and Other Reptiles of Southern Africa. 3rd ed. Fully Revised and Updated to Include 83 New Species. Ralph Curtis Books (Sanibel Island, Florida), 399 pp.
  • Fitzsimons, V.F.M.; Brain, C.K. 1958. A Short account of the Reptiles of the Kalahari Gemsbok National Park. Koedoe, 1(1): 99-104 - get paper here
  • Fitzsimons, Vivian Frederick Maynard 1937. Notes on the reptiles and amphibians collected and described from South Africa by Andrew Smith. Annals Transvaal Museum 17: 259-274. - get paper here
  • Fitzsimons,V.F.M. 1941. Descriptions of some new lizards from South Africa and a frog from southern Rhodesia. Annals Transvaal Mus. 20 (3): 273-281 - get paper here
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • Hewitt, J. 1927. Further descriptions of reptiles and batrachians from South Africa. Rec. Albany Mus. (Grahamstown) 3 (5): 371-415
  • Hewitt, J. 1935. Some new forms of batrachians and reptiles from South Africa. Rec. Albany Mus. 4: 283-357
  • Lamb, T. & A. M. BAUER 2000. Relationships of the Pachydactylus rugosus group of geckos (Reptilia: Squamata: Gekkonidae). African Zoology 35 (1): 55-67 - get paper here
  • Loveridge, A. 1947. Revision of the African lizards of the family Gekkondiae. Bull. Mus. Comp. Zool. Harvard 98: 1-469 - get paper here
  • McLachlan G R 1979. The taxonomy of Pachydactylus rugosus. J. Herp. Assoc. Africa (No. 21): 4-7 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Schleicher, Alfred 2020. Reptiles of Namibia. Kuiseb Publishers, Windhoek, Namibia, 271 pp.
  • Smith, A. 1849. Illustrations of the Zoology of South Africa. 3 (Reptiles). Smith, Elder, and Co., London [facsimile published by Winchester Press, Johannesburg, 1977] - get paper here
  • Thominot, A. 1889. Observations sur quelques reptiles et batraciens de la collection du Muséum d’Histoire Naturelle de Paris. Bulletin de la Société Philomathique de Paris (8) 1:21—30 - get paper here
 
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