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Philodryas psammophidea GÜNTHER, 1872

IUCN Red List - Philodryas psammophidea - Least Concern, LC

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesGünther's Green Racer 
SynonymPhilodryas psammophideus GÜNTHER 1872: 23
Dirrhox lativittatus COPE 1887: 58
Philodryas bolivianus BOULENGER 1896: 132
Philodryas psammophideus — BOULENGER 1896 (3): 132
Philodryas Borellii PERACCA 1897
Liophis trifasciatus WERNER 1899: 114
Liophis bolivianus WERNER 1909: 222
Philodryas lineatus WERNER 1909
Philodryas werneri MÜLLER 1925
Philodryas pallidus WERNER 1926
Chlorosoma psammophideum — SERIÉ 1936
Philodryas psammophideus — PETERS & OREJAS-MIRANDA 1970: 244
Dirrhox lattivittatus — MALNATE 1971: 368 (in error)
Philodryas psammophideus — FREIBERG 1982: 150
Philodryas psammophideus psammophideus — CEI 1993
Philodryas bolivianus — CEI 1993
Philodryas psammophideus — LEYNAUD & BUCHER 1999: 31
Philodryas psammophidea — BERNILS et al. 2007
Philodryas psammophidea — GRAZZIOTIN et al. 2012
Philodryas psammophidea — WALLACH et al. 2014: 552
Philodryas psammmophidea — RIVAS et al. 2018 (in error) 
DistributionWC Brazil, E Bolivia, Paraguay,
Argentina (Rio Negro, Neuquén, Mendoza, San Juan, La Rioja, Catamarca, Tucumán, Salta, Jujuy, Chaco, Formosa, Santiago del Estero, Córdoba, Santa Fe, San Luis, La Pampa, Corrientes, Chubut)

Type locality: Tucumán, Argentina

boliviana: Bolivia; Type locality: Charobamba, Bolivia

werneri: Type locality: “Sierra de Curumalan, Ost-Argentinien” [Buenos Aires prov., Argentina]  
Reproductionoviparous 
TypesHolotype: BMNH 1946.1.2.79, a 686 mm female.
Type: MSNTO R226, R550 (P. borellii)
Holotype: ZSM 43/1974 (originally ZMB 16860), juvenile [werneri] 
Diagnosis 
CommentSynonymy after PETERS & OREJAS-MIRANDA 1970 and CEI 1993 (but see note for P. borelli), partly following AMARAL 1926. Philodryas bolivianus was considered a s synonym of Philodryas psammophideus by PETERS & OREJAS-MIRANDA 1970, but not by CEI 1993.

Distribution: This species has been recorded from Uruguay inn the past but according to CARREIRA et al. (2005) there are no reliable records from that country. 
Etymology 
References
  • Abalos, J.W., Baez, E.C., & Nader, R. 1964. Serpientes de Santiago del Estero. Acta Zoologica Lilloana 20: 211-283.
  • Amaral,A. do 1926. Notas de ophiologia. I. Sobre a invalidez de um genero e algumas especies de ophidios Sul-Americanos. Rev. Mus. Paulista 14: 17-33
  • Boulenger, G.A. 1896. Catalogue of the snakes in the British Museum, Vol. 3. London (Taylor & Francis), xiv + 727 pp. - get paper here
  • Cacciali, Pier; Norman J. Scott, Aida Luz Aquino Ortíz, Lee A. Fitzgerald, and Paul Smith 2016. The Reptiles of Paraguay: Literature, Distribution, and an Annotated Taxonomic Checklist SPECIAL PUBLICATION OF THE MUSEUM OF SOUTHWESTERN BIOLOGY, NUMBER 11: 1–373 - get paper here
  • Cei, J. M. 1993. Reptiles del noroeste, nordeste y este de la Argentina. Museo Regionale Sci. Naturale Torino, Monografie 14: 1-949
  • Cope, E.D. 1887. Synopsis of the Batrachia and Reptilia obtained by H. H. Smith in the Province of Mato Grosso, Brazil. Proc. Amer. Philos. Soc. 24: 44-60 - get paper here
  • Franca, F.G.R.; Daniel O. Mesquita, Cristiano C. Nogueira, and Alexandre F. B. Araújo 2008. Phylogeny and Ecology Determine Morphological Structure in a Snake Assemblage in the Central Brazilian Cerrado. Copeia 2008 (1): 23-38 - get paper here
  • Freiberg, M.A. 1982. Snakes of South America. T.F.H. Publications, Neptune City, NJ, 189 pp.
  • Gonzales, Lucindo; Arturo Muñoz y Edson Cortez 2006. PRIMER REPORTE SOBRE LA HERPETOFAUNA DE LA RESERVA NATURAL “EL CORBALÁN”, TARIJA, BOLIVIA [FIRST REPORT ON THE HERPETOFAUNA OF “EL CORBALAN” NATURAL RESERVE, TARIJA, BOLIVIA]. Kempffiana 2 (1): 72-94 - get paper here
  • Grazziotin, Felipe G.; Hussam Zaher, Robert W. Murphy, Gustavo Scrocchi, Marco A. Benavides, Ya-Ping Zhang and Sandro L. Bonatto 2012. Molecular phylogeny of the New World Dipsadidae (Serpentes: Colubroidea): a reappraisal. Cladistics DOI: 10.1111/j.1096-0031.2012.00393.x - get paper here
  • Günther,A. 1872. Seventh account of new species of snakes in the collection of the British Museum. Ann. Mag. Nat. Hist. (4) 9: 13-37 - get paper here
  • Hohmeister, A. 1999. Bemerkungen zur Biologie und Pflege südamerikanischer Strauchnattern der Gattung Philodryas Wagler, 1830 im Terrarium. Sauria 21 (2): 9-21 - get paper here
  • Kacoliris F.P.; Berkunsky I. & Williams J. 2006. Herpetofauna of Impenetrable, Argentinean Great Chaco. Phyllomedusa 5 (2): 149-158 - get paper here
  • Leynaud, Gerardo C. and Enrique H. Bucher 1999. La fauna de serpientes del Chaco Sudamericano: diversidad, distribución geografica y estado de conservación. Academia Nacional de Ciencias Miscelanea (98):1-46
  • Malnate, Edmund V. 1971. A catalogue of primary types in the Herpetological Collections of the Academy of Natural Sciences, Philadelphia (ANSP). Proccedings of the Academy of Natural Sciences of Philadelphia 123 (9): 345-375 - get paper here
  • Mattison, Chris 2007. The New Encyclopedia of Snakes. Princeton University Press
  • Minoli I, Morando M, Avila LJ 2015. Reptiles of Chubut province, Argentina: richness, diversity, conservation status and geographic distribution maps. ZooKeys 498: 103-126. doi: 10.3897/zookeys.498.7476 - get paper here
  • Pereyra, Laura Cecilia and Mauricio Sebastián Akmentins. 2013. A case of predation of Chacophrys pierottii (Anura: Ceratophryidae) by Philodryas psammophidea (Squamata: Dipsadinae) in the chacoan saltflats of central Argentina. Herpetology Notes 6: 245-246. - get paper here
  • Quinteros-Muñoz, Oliver; Peñaranda, Diego A.; Navarro, Freddy 2010. Rodent consumption by Philodryas psammophidea (Serpentes: Colubridae), from the inter-andean dry valleys of central Bolivia. Cuadernos de Herpetología 24 (2) - get paper here
  • Ramalho, Werther Pereira; Jefferson Rocha Silva, Philip Teles Soares, Denes Ferraz, Filipe Viegas Arruda, Vitor Hugo Mendonça do Prado 2018. The anurans and squamates of a peri-urban Cerrado remnant in the State of Goiás, Central Brazil Herpetology Notes 11: 573-583 - get paper here
  • Rivas, Luis Rolando; Mendoza-Miranda, Patricia; Muñoz-Saravia, Arturo 2018. Lista preliminar de la herpetofauna del Parque Nacional Torotoro, Potosí, Bolivia . Pp. 6 Cuad. herpetol. 32 (1): 41-46 - get paper here
  • Scrocchi, G.J.; Moreta, J.C. & Kretzschmar, S. 2006. Serpientes del Noroeste Argento [Jujuy, Salta, Tucumán, Catamarca, La Rioja, Santiago del Estero]. Fundación Miguel Lillo, Tucumán, 178 pp.
  • Scrocchi, Gustavo J.; Cristian S. Abdala,Javier Nori, Hussam Zaher 2010. Reptiles de la provincia de Río Negro, Argentina Fondo Ed. Rionegrino, 249 pp.
  • Thomas, R. A. 1976. A revision of the South American colubrid snake genus Philodryas Wagler, 1830. PhD Thesis, Texas A & M Univ.: 378 pp.
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Werner, F. 1899. Beschreibungen einiger neuer Schlangen und Batrachier. Zool. Anz. 22: 114-117 - get paper here
  • Werner, FRANZ 1901. Reptilien und Batrachier aus Peru und Bolivien. Abhandl. Ber. Zool. Anthrop. Mus. Dresden, 9 (2): 1-14 [1900]
  • Williams,J.D. & Francini,F. 1991. A checklist of the Argentine snakes. Boll. Mus. Reg. Sci. Nat. Torino 9 (1): 55-90
  • Zaher, H.; Scrocchi, G. & Masiero, R. 2008. Rediscovery and redescription of the type of Philodryas laticeps Werner, 1900 and the taxonomic status of P. oligolepis Gomes, 1921 (Serpentes, Colubridae). Zootaxa 1940: 25–40 - get paper here
 
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