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Pholidoscelis auberi (COCTEAU, 1838)

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Higher TaxaTeiidae, Teiinae, Gymnophthalmoidea, Sauria, Squamata (lizards) 
SubspeciesPholidoscelis auberi auberi (COCTEAU 1838)
Pholidoscelis auberi abducta (SCHWARTZ 1970)
Pholidoscelis auberi atrothorax (SCHWARTZ 1970)
Pholidoscelis auberi behringensis (LEE & SCHWARTZ 1985)
Pholidoscelis auberi bilateralis (MCCOY 1970)
Pholidoscelis auberi cacuminis (SCHWARTZ 1970)
Pholidoscelis auberi citra (SCHWARTZ 1970)
Pholidoscelis auberi denticola (SCHWARTZ 1970)
Pholidoscelis auberi extorris (SCHWARTZ 1970)
Pholidoscelis auberi extraria (SCHWARTZ 1970)
Pholidoscelis auberi felis (MCCOY 1970)
Pholidoscelis auberi focalis (MCCOY 1970)
Pholidoscelis auberi galbiceps (SCHWARTZ 1970)
Pholidoscelis auberi garridoi (SCHWARTZ 1970)
Pholidoscelis auberi gemmea (SCHWARTZ 1970)
Pholidoscelis auberi granti (SCHWARTZ 1970)
Pholidoscelis auberi hardyi (SCHWARTZ 1970)
Pholidoscelis auberi kingi (MCCOY 1970)
Pholidoscelis auberi llanensis (SCHWARTZ 1970)
Pholidoscelis auberi marcida (SCHWARTZ 1970)
Pholidoscelis auberi multilineata (MCCOY 1970)
Pholidoscelis auberi nigriventris (GALI & GARRIDO 1987)
Pholidoscelis auberi obsoleta (MCCOY in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi orlandoi (SCHWARTZ & MCCOY 1975)
Pholidoscelis auberi parvinsulae (LEE & SCHWARTZ 1985)
Pholidoscelis auberi paulsoni (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi peradusta (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi procer (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi pullata (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi richmnondi (MCCOY in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi sabulicolor (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi sanfelipensis (GARRIDO 1975)
Pholidoscelis auberi schwartzi (GALI & GARRIDO 1987)
Pholidoscelis auberi secta (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi sideroxylon (LEE & SCHWARTZ 1985)
Pholidoscelis auberi sublesta (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi thoracica (COPE 1863)
Pholidoscelis auberi ustulata (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi vulturnus (LEE & SCHWARTZ 1985)
Pholidoscelis auberi zugi (SCHWARTZ in SCHWARTZ & MCCOY 1970) 
Common NamesAuber's Ameiva 
SynonymAmeiva auberi COCTEAU 1838: 51
Ameiva auberi — DUMÉRIL & BIBRON 1839: 111
Ameiva auberi — SCHWARTZ & HENDERSON 1988: 53
Ameiva auberi — SCHWARTZ & HENDERSON 1991: 182
Ameiva auberi — HARVEY et al. 2012
Ameiva auberi — RODRIGUEZ-SCHETTINO et al. 2013: 21
Pholidoscelis auberi — GOICOECHEA et al. 2016

Pholidoscelis auberi orlandoi (SCHWARTZ & MCCOY 1975)
Ameiva auberi festiva GARRIDO 1975
Ameiva auberi orlandoi SCHWARTZ & MCCOY 1975 (nom. subst.)
Ameiva auberi orlandoi — SCHWARTZ & HENDERSON 1988: 57
Ameiva auberi orlandoi — ESTRADA 1998
Pholidoscelis auberi orlandoi — GOICOECHEA et al. 2016 (by implication)

Pholidoscelis auberi sabulicolor (SCHWARTZ 1970)
Ameiva auberi sabulicolor SCHWARTZ 1970
Ameiva auberi sabulicolor — SCHWARTZ & HENDERSON 1988: 58

Pholidoscelis auberi thoracica (COPE 1862)
Ameiva thoracica COPE 1862: 64
Ameira [sic] thoracica — BARBOUR 1904: 59
Ameiva thoracica — BARBOUR & SHREVE 1935: 361
Ameiva auberi thoracica — SCHWARTZ & HENDERSON 1988: 59
Pholidoscelis auberi thoracica — GOICOECHEA et al. 2016 (by implication)

Ameiva auberi — SCHWARTZ & HENDERSON 1988: 53

Pholidoscelis auberi abducta (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Ameiva auberi abducta SCHWARTZ 1970: 70
Ameiva auberi abducta — SCHWARTZ & HENDERSON 1988: 53

Pholidoscelis auberi atrothorax (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Ameiva auberi atrothorax SCHWARTZ 1970: 79
Ameiva auberi atrothorax — SCHWARTZ & HENDERSON 1988: 53

Pholidoscelis auberi behringensis (LEE & SCHWARTZ 1985)
Ameiva auberi behringensis LEE & SCHWARTZ 1985: 13
Ameiva auberi behringensis — SCHWARTZ & HENDERSON 1988: 54

Pholidoscelis auberi bilateralis (MCCOY 1970)
Ameiva auberi bilateralis MCCOY 1970: 79
Ameiva auberi bilateralis — SCHWARTZ & HENDERSON 1988: 54

Pholidoscelis auberi cacuminis (SCHWARTZ 1970)
Ameiva auberi cacuminis SCHWARTZ in SCHWARTZ & MCCOY 1970: 56
Ameiva auberi cacuminis — SCHWARTZ & HENDERSON 1988: 54

Pholidoscelis auberi citra (SCHWARTZ 1970)
Ameiva auberi citra SCHWARTZ in SCHWARTZ & MCCOY 1970: 89
Ameiva auberi citra — SCHWARTZ & HENDERSON 1988: 54

Pholidoscelis auberi denticola (SCHWARTZ 1970)
Ameiva auberi denticola SCHWARTZ in SCHWARTZ & MCCOY 1970: 57
Ameiva auberi denticola — SCHWARTZ & HENDERSON 1988: 54

Pholidoscelis auberi extorris (SCHWARTZ 1970)
Ameiva auberi extorris SCHWARTZ in SCHWARTZ & MCCOY 1970: 76
Ameiva auberi extorris — SCHWARTZ & HENDERSON 1988: 54

Pholidoscelis auberi extraria (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Ameiva auberi extraria SCHWARTZ in SCHWARTZ & MCCOY 1970: 73
Ameiva auberi extraria — SCHWARTZ & HENDERSON 1988: 54

Pholidoscelis auberi felis (MCCOY 1970)
Ameiva auberi felis SCHWARTZ in SCHWARTZ & MCCOY 1970: 128
Ameiva auberi felis — SCHWARTZ & HENDERSON 1988: 55

Pholidoscelis auberi focalis (MCCOY 1970)
Ameiva auberi focalis SCHWARTZ in SCHWARTZ & MCCOY 1970: 137
Ameiva auberi focalis — SCHWARTZ & HENDERSON 1988: 55

Pholidoscelis auberi galbiceps (SCHWARTZ 1970)
Ameiva auberi galbiceps SCHWARTZ in SCHWARTZ & MCCOY 1970: 111
Ameiva auberi galbiceps — SCHWARTZ & HENDERSON 1988: 55

Pholidoscelis auberi garridoi (SCHWARTZ 1970)
Ameiva auberi garridoi SCHWARTZ in SCHWARTZ & MCCOY 1970: 77
Ameiva auberi garridoi — SCHWARTZ & HENDERSON 1988: 54

Pholidoscelis auberi gemmea (SCHWARTZ 1970)
Ameiva auberi gemmea SCHWARTZ in SCHWARTZ & MCCOY 1970: 71
Ameiva auberi gemmea — SCHWARTZ & HENDERSON 1988: 55

Pholidoscelis auberi granti (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Ameiva auberi granti SCHWARTZ in SCHWARTZ & MCCOY 1970: 91
Ameiva auberi granti — SCHWARTZ & HENDERSON 1988: 55 
DistributionCuba, Isla de la Juventud, Cat Island, Grand Bahama Bank

auberi: western Cuba, east to Matanzas and La Habana provinces. Type locality: Cuba; restricted to the cicinity of La Habana, La Habana Province, Cuba by SCHWARTZ 1970.

abducta: Cuba, distal half of the Peninsula de Hicacos. Type locality: Punta Hicacos, Matanzas Province, Cuba.

atrothorax: South-central Cuba, in Cienfuegos, Sancti Spiritus, and Villa Clara provinces. Type locality: Finca Morales, 8 mi. NW Trinidad, Sancti Spiritus Province, Cuba.

behringensis: Bahama Is.: Andros I. Type locality: Behring Point, Andros Island, Bahama Islands.

bilateralis: Bahama Is.: Ragged Is. Type locality: South end of Great Ragged Island, Bahama Islands.

cacuminis: Cuba, extreme distal Peninsula de Guanahacabibes. Type locality: Ensenada de Caj6n, Pinar del Rio Province, Cuba.

citra: Cuba, northern Camagüey and Las Tunas provinces. Type locality: 2 mi. W Playa Santa Lucia, Camagiiey Province, Cuba

denticola: Cuba, proximal portion of Peninsula de Guanahacabibes. Type locality: North shore, Ensenada de Corrientes, Pinar del Rio Province, Cuba.

extorris: Cuba, Archipiélago de Sabana-Camagüey. Type locality: Cayuelo de la Vela, Villa Clara Province, Cuba.

extraria: Cuba, Archipiélago de Sabana-Camagüey; material from Cayo Carenero intergradient with A. a. extorris. Type locality: Cayo Bahia de Cadiz, Villa Clara Province, Cuba.

felis: Bahama Is.: Cat I. Type locality: The Bight, Cat Island, Bahama Islands.

focalis: Bahama Is.: northern Exuma Cays. Type locality: Ship Channel Cay, Exuma Cays, Bahama Islands.

galbiceps: Cuba, Jardines de la Reina. Type locality: Southernmost point of large cay, 3 km NW Cayo Cachiboca, Laberinto de las Doce Leguas, Camagiiey Province, Cuba

garridoi: Cuba, northern Villa Clara Province. Type locality: Cuatro Bocas, Sagua la Grande, Villa Clara Province, Cuba.

gemmea: Cuba, mouth of Rio de Sierra Morena, Villa Clara Province; perhaps at Baños de Elguea, Salinas de Bidos, and on Cayo Cinco Leguas, Matanzas Province. Type locality: Mouth of Rio de Sierra Morena, near Playa Ganuza, Villa Clara Province, Cuba.

granti: Cuba, northern Holguin and Guantánamo provinces. Type locality: Baracoa, east side, Bahia de Miel, Guantanamo Province, Cuba.

hardyi: Cuba, southeastern coast in Granma and Santiago de Cuba provinces, and probably on all the northern slopes of the Sierra Maestra. Type locality: Ocujal, Santiago de Cuba Province, Cuba.

kingi: Bahama Is.: Andros I., Gibson Cay, Big Wood Cay; intergrades with A. a. vulturnus on Mangrove Cay. Type locality: Gibson Cay, mouth of South Bight, Andros Island, Bahama Islands.

llanensis: Cuba, serpentine savannas in Camagüey Province. Type locality: Just south of west end of Sierra de Cubitas, Camagiiey Province, Cuba.

marcida: Isla de la Juventud, south of the Ciénaga de Lanier. Type locality: Jacksonville, Isla de la Juventud, Cuba.

multilineata: Bahama Is.: Berry Is. Type locality: Bond's Cay, Berry Islands, Bahama Islands.

nigriventris: Isla de la Juventud, NE of the Ciénaga de Lanier.
Type locality: Outskirts of Cayo Piedras, Isla de la Juventud, Cuba.

obsoleta: Bahama Is.: Long I., Exuma Cays. Type locality: Adderly's, Long Island, Bahama Islands.

orlandoi: Cuba, Archipiélago de Sabana-Camagüey; also at Buena Vista, Ciego de Avila Province. Type locality: Cayo Santa Maria, Archipielago de Sabana-Camagliey, Villa Clara Province, Cuba.

parvinsulae: Bahama Is.: Green Cay. Type locality: Green Cay, Bahama Islands.

paulsoni: Cuba, southern Pinar del Rio Province. Type locality: 1 km N Las Canas, Pinar del Rio Province, Cuba.

peradusta: Cuba, from Juragmá to Calimete, thence to about 25 km B Playa Girón, Peninsula de Zapata, and inland to northern Cienfuegos Province. Type locality: Juragua, Cienfuegos Province, Cuba.

procer: Cuba, throughout much of Pinar del Rio Province. Type locality: San Vicente, Pinar del Rio Province, Cuba.

pullata. Cuba, northern Matanzas Province. Type locality: 13 km NE Matanzas, Matanzas Province, Cuba.

richmondi: Bahama Is.: Bimini Is. Type locality: Near Lyons, North Bimini Island, Bahama Islands.

sabulicolor: Cuba, southeastern coast. Type locality: 2.8 mi. E Imias, Guantanamo Province, Cuba.

sanfelipensis: Cuba, Cayos de San Felipe. Type locality: Cayo
Real, Cayos de San Felipe, Pinar del Rio Province, Cuba.

schwartzi: Cuba, northeastern Pinar del Rio Province. Type locality: Henequenera de Mariel, Zona Mosquito, Pinar del Rio Province, Cuba.

secta: Isla de la Juventud, north of the Ciénaga de Lanier, except range of A. a. nigriventris. Type locality: Playa de Rocas, between Bibijagua and Ju.caro, Isla de la Juven- tud, Cuba.

sideroxylon: Bahama Is.: Andros I. Type locality: 4.8 km N Mastic Point, Andros Island, Bahama Islands.

sublesta: Cuba, Archipiélago de Sabana-Camagüey. Type locality: Playa Bonita, east end, Cayo Sabinal, Camagiiey Province, Cuba.

thoracica: Bahama Is.: New Providence I., Rose I., Eleuthera I., Eleuthera Cays, Little San Salvador I. Type locality: New Providence Island, Bahama Islands.

ustulata: Cuba, southeastern coast between Bahia de Santiago and Bahia de GuantAnamo, and inland north of Santiago de Cuba. Type locality: 7.8 mi. E Siboney, Santiago de Cuba Province, Cuba.

vulturnus: Bahama Is.: Andros I. Type locality: Marsh Bay, Andros Island, Bahama Islands.

zugi: Cuba, Archipiélago de los Canarreos, and Peninsula de Zapata. Type locality: Cayo Largo del Sur, Archipielago de los Canarreos, Isla de la Juventud, Cuba.

INCERTAE SEDIS: Material from Herradura, Pinar del Rio Province; Cayo Macho de Tierra, about 10 mi. SE Casilda; Puerto Padre and Gibara, Holguin Province; Sierra de Jatibonico, Sabana San Felipe, Arroyo Blanco, Isla de Turiguanó is unassigned subspecifically.  
Reproductionoviparous 
TypesSyntypes: MNHN 1112, MNHN 2647, MNHN 1788, MNHN 4178 (see Schwartz, 1970a) (auberi)
Holotype: ANSP 9158 [thoracica]
Holotype: AMNH 96331 [abducta]
Holotype: AMNH 78035 [atrothorax]
Holotype: CM 60540 [behringensis]
Holotype: CM 40985 [bilateralis]
Holotype: AMNH 83028 [cacuminis]
Holotype: AMNH 79202 [denticola]
Holotype: IZ 52 [extorris]
Holotype: AMNH 82982 [extraria]
Holotype: CM 20440 [felis]
Holotype: CM 41147 [focalis]
Holotype: AMNH 78058 [galbiceps]
Holotype: IZ 96 [garridoi]
Holotype: AMNH 82972 [gemmea]
Holotype: AMNH 83784 [granti]
Holotype: USNM 138468 [hardyi]
Holotype: CM 40915 [kingi]
Holotype: AMNH 82991 [marcida]
Holotype: CM 41196 [multilineata]
Holotype: IZ 5778 [nigriventris]
Holotype: CM 43976 [obsoleta]
Holotype: IZ 3427 [orlandoi]
Holotype: CM 60542 [parvinsulae]
Holotype: AMNH 83012 [paulsoni]
Holotype: IZ 85 [peradusta]
Holotype: AMNH 78390 [procer]
Holotype: AMNH 82953 [pullata]
Holotype: CM 34140 [richmnondi]
Holotype: AMNH 83941 [sabulicolor]
Holotype: IZ 2987 [sanfelipensis]
Holotype: IZ 4389 [schwartzi]
Holotype: AMNH 82997 [secta]
Holotype: CM 60539 [sideroxylon]
Holotype: AMNH 96393 [sublesta]
Holotype: ANSP 9158 [thoracica]
Holotype: AMNH 83778 [ustulata]
Holotype: CM 60541 [vulturnus]
Holotype: AMNH 82003 [zugi]
Holotype: MCZ 77049, adult male; paratypes: MCZ, AMNH, KU, USNM [citra]
Holotype: AMNH 96375. [citra]
Holotype: MCZ 59321. [llanensis] 
CommentIllustrations: Barbour and Ramsden, 1919; Schwartz and Henderson, 1985.

Subspecies: RODRIGUEZ-SCHETTINO et al. 2013 do not distinguish between subspecies, but see Schwartz 1970: 113 for a map of the Cuban subspecies. 
EtymologyNamed after Professor Pedro Alejandro Auber (1786-1843), a Cuban botanist and naturalist of French origin (née Pierre Alexandre Auber).

Pholidoscelis auberi zugi was named in honour of Professor Dr. George Zug, National Museum of Natural History, Smithsonian Institution (Washington DC, USA). For biographical details see Mitchell 2018. 
References
  • Barbour, T. 1904. Batrachia and Reptilia from the Bahamas. Bull. Mus. Comp. Zool. Harvard 46: 55—61 - get paper here
  • Barbour, T. and B. Shreve. 1935. Concerning some Bahamian reptiles, with notes on the fauna. Proc. Boston Soc. Nat. Hist. 40: 347-365.
  • Barbour, T. and C. T. Ramsden. 1919. The herpetology of Cuba. Mem. Mus. Comp. Zool., Harvard Coll. 47: 69-213. - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Cocteau, J.-T. and G. Bibron 1838. Reptiles, in R. de la Sagra. Historia Física, Política y Natural de la Isla de Cuba, Tomo IV. Arthus Bertrand, Paris, 142 pp. [1838-1843 - index on page 143] - get paper here
  • Cope, E.D. 1862. Synopsis of the species of Holcosus and Ameiva, with diagnoses of new West Indian and South American Colubridae. Proc. Acad. Nat. Sci. Philadelphia 14: 60-82. - get paper here
  • Duméril, A. M. C. and G. Bibron. 1839. Erpétologie Générale on Histoire Naturelle Complète des Reptiles. Vol.5. Roret/Fain et Thunot, Paris, 871 pp. - get paper here
  • Estrada, A.R. 1998. Anfibios y Reptiles Encontrados Durante 1988 y 1989 en Cayo Paredón Grande, Archipiélago de Sabana-Camagüey, Cuba. Carib. J. Sci. 34 (1-2): 106-112
  • Franz, Richard;Dodd, C. Kenneth, Jr 1994. Ameiva auberi obsoleta (Exuma Islands ameiva). Bahamas: Great Bahama Bank. Herpetological Review 25 (4): 163-164 - get paper here
  • Gali, F. and Garrido 1987. Two new subspecies of Ameiva auberi (Reptilia: Teiidae) from Cuba. Carib J. Sci. 22 (3-4): 165-173 [1986]
  • Garrido, O. H. 1975. Nuevos reptiles del archipielago cubano. Poeyana 141:1-58.
  • Goicoechea, N., Frost, D. R., De la Riva, I., Pellegrino, K. C. M., Sites, J., Rodrigues, M. T. and Padial, J. M. 2016. Molecular systematics of teioid lizards (Teioidea/Gymnophthalmoidea: Squamata) based on the analysis of 48 loci under tree-alignment and similarity-alignment. Cladistics, doi: 10.1111/cla.12150 - get paper here
  • Hower, Lindsey M., and S. Blair Hedges 2003. Molecular phylogeny and biogeography of West Indian Teiid lizards of the genus Ameiva. Carib. J. Sci. 39 (3):298-306. - get paper here
  • Lee, Daniel K.;Schwartz, Albert 1985. Four new subspecies of Ameiva auberi (Sauria, Teiidae) from the Bahama Islands. Annals of the Carnegie Museum 54 (2): 11-21
  • Mitchell, Joseph C. 2018. George Robert Zug. Copeia 106 (2): 388-395 - get paper here
  • Rodríguez-Cabrera, Tomás M.; Alejandro M. Rodríguez González, and Javier Torres 2018. Herpetofauna of Cayo Carenas, Cienfuegos Bay, south-central Cuba IRCF Reptiles & Amphibians 25 (1): 13–19 - get paper here
  • Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96 - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Schwartz, A. & McCoy, C.J. 1970. A systematic review of Ameiva auberi Cocteau (Reptilia, Teiidae) in Cuba and the Bahamas. I. The Cuban subspecies. II: The Bahamian subspecies. III. Discussion. Annals of the Carnegie Museum 41(4): 45-168.
  • Schwartz, A. and Henderson, R.W. 1985. A guide to the identification of the amphibians and reptiles of the West Indies exclusive of Hispaniola. Milwaukee Public Mus., 165 pp.
  • Schwartz, A. and McCoy, C.J. 1975. A New Name for a Cuban Ameiva (Reptilia: Teiidae) Herpetologica 31(2): 240 - get paper here
  • Tucker, Derek B.; | Stephen Blair Hedges | Guarino R. Colli | Robert Alexander Pyron | Jack W. Sites Jr 2017. Genomic timetree and historical biogeography of Caribbean island ameiva lizards (Pholidoscelis: Teiidae) Ecology and Evolution 7:7080–7090.
  • Varona, Luis S.;Garrido, Orlando H. 1970. Vertebrados de los Cayos de San Felipe, Cuba, Inclyendo una nueva especie de Jutía. Poeyana (75): 1-26
 
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