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Pholidoscelis auberi (COCTEAU, 1838)

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Higher TaxaTeiidae, Teiinae, Gymnophthalmoidea, Sauria, Squamata (lizards) 
SubspeciesPholidoscelis auberi auberi (COCTEAU 1838)
Pholidoscelis auberi abducta (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi atrothorax (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi behringensis (LEE & SCHWARTZ 1985)
Pholidoscelis auberi bilateralis (MCCOY 1970)
Pholidoscelis auberi cacuminis (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi citra (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi denticola (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi extorris (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi extraria (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi felis (MCCOY in SCHWARTZ & MCCOY)
Pholidoscelis auberi focalis (MCCOY in SCHWARTZ & MCCOY)
Pholidoscelis auberi galbiceps (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi garridoi (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi gemmea (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi granti (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi hardyi (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi kingi (MCCOY in SCHWARTZ & MCCOY)
Pholidoscelis auberi llanensis (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi marcida (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi multilineata (MCCOY in SCHWARTZ & MCCOY)
Pholidoscelis auberi nigriventris (GALI & GARRIDO 1987)
Pholidoscelis auberi obsoleta (MCCOY in SCHWARTZ & MCCOY)
Pholidoscelis auberi orlandoi (SCHWARTZ & MCCOY 1975)
Pholidoscelis auberi parvinsulae (LEE & SCHWARTZ 1985)
Pholidoscelis auberi paulsoni (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi peradusta (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi procer (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi pullata (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi richmnondi (MCCOY in SCHWARTZ & MCCOY)
Pholidoscelis auberi sabulicolor (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi sanfelipensis (GARRIDO 1975)
Pholidoscelis auberi schwartzi (GALI & GARRIDO 1987)
Pholidoscelis auberi secta (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi sideroxylon (LEE & SCHWARTZ 1985)
Pholidoscelis auberi sublesta (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi thoracica (COPE 1863)
Pholidoscelis auberi ustulata (SCHWARTZ in SCHWARTZ & MCCOY 1970)
Pholidoscelis auberi vulturnus (LEE & SCHWARTZ 1985)
Pholidoscelis auberi zugi (SCHWARTZ in SCHWARTZ & MCCOY 1970) 
Common NamesAuber's Ameiva 
SynonymAmeiva auberi COCTEAU 1838: 51
Ameiva auberi — DUMÉRIL & BIBRON 1839: 111
Ameiva auberi — SCHWARTZ & HENDERSON 1988: 53
Ameiva auberi — SCHWARTZ & HENDERSON 1991: 182
Ameiva auberi — HARVEY et al. 2012
Ameiva auberi — RODRIGUEZ-SCHETTINO et al. 2013: 21
Pholidoscelis auberi — GOICOECHEA et al. 2016

Pholidoscelis auberi orlandoi (SCHWARTZ & MCCOY 1975)
Ameiva auberi festiva GARRIDO 1975
Ameiva auberi orlandoi SCHWARTZ & MCCOY 1975 (nom. subst.)
Ameiva auberi orlandoi — SCHWARTZ & HENDERSON 1988: 57
Ameiva auberi orlandoi — ESTRADA 1998
Pholidoscelis auberi orlandoi — GOICOECHEA et al. 2016 (by implication)

Pholidoscelis auberi sabulicolor (SCHWARTZ 1970)
Ameiva auberi sabulicolor SCHWARTZ 1970
Ameiva auberi sabulicolor — SCHWARTZ & HENDERSON 1988: 58

Pholidoscelis auberi thoracica (COPE 1862)
Ameiva thoracica COPE 1862: 64
Ameira [sic] thoracica — BARBOUR 1904: 59
Ameiva thoracica — BARBOUR & SHREVE 1935: 361
Ameiva auberi thoracica — SCHWARTZ & HENDERSON 1988: 59
Pholidoscelis auberi thoracica — GOICOECHEA et al. 2016 (by implication) 
DistributionCuba, Isla de la Juventud, Cat Island, Grand Bahama Bank

auberi: western Cuba, east to Matanzas and La Habana provinces. Type locality: Cuba; restricted to the cicinity of La Habana, La Habana Province, Cuba by SCHWARTZ 1970.

abducta: Cuba, distal half of the Peninsula de Hicacos. Type locality: Punta Hicacos, Matanzas Province, Cuba.

atrothorax: South-central Cuba, in Cienfuegos, Sancti Spiritus, and Villa Clara provinces. Type locality: Finca Morales, 8 mi. NW Trinidad, Sancti Spiritus Province, Cuba.

behringensis: Bahama Is.: Andros I. Type locality: Behring Point, Andros Island, Bahama Islands.

bilateralis: Bahama Is.: Ragged Is. Type locality: South end of Great Ragged Island, Bahama Islands.

cacuminis: Cuba, extreme distal Peninsula de Guanahacabibes. Type locality: Ensenada de Caj6n, Pinar del Rio Province, Cuba.

citra: Cuba, northern Camagüey and Las Tunas provinces. Type locality: 2 mi. W Playa Santa Lucia, Camagiiey Province, Cuba

denticola: Cuba, proximal portion of Peninsula de Guanahacabibes. Type locality: North shore, Ensenada de Corrientes, Pinar del Rio Province, Cuba.

extorris: Cuba, Archipiélago de Sabana-Camagüey. Type locality: Cayuelo de la Vela, Villa Clara Province, Cuba.

extraria: Cuba, Archipiélago de Sabana-Camagüey; material from Cayo Carenero intergradient with A. a. extorris. Type locality: Cayo Bahia de Cadiz, Villa Clara Province, Cuba.

felis: Bahama Is.: Cat I. Type locality: The Bight, Cat Island, Bahama Islands.

focalis: Bahama Is.: northern Exuma Cays. Type locality: Ship Channel Cay, Exuma Cays, Bahama Islands.

galbiceps: Cuba, Jardines de la Reina. Type locality: Southernmost point of large cay, 3 km NW Cayo Cachiboca, Laberinto de las Doce Leguas, Camagiiey Province, Cuba

garridoi: Cuba, northern Villa Clara Province. Type-locality: Cuatro Bocas, Sagua la Grande, Villa Clara Province, Cuba.

gemmea: Cuba, mouth of Rio de Sierra Morena, Villa Clara Province; perhaps at Baños de Elguea, Salinas de Bidos, and on Cayo Cinco Leguas, Matanzas Province. Type-locality: Mouth of Rio de Sierra Morena, near Playa Ganuza, Villa Clara Province, Cuba.

granti: Cuba, northern Holguin and Guantánamo provinces. Type locality: Baracoa, east side, Bahia de Miel, Guantanamo Province, Cuba.

hardyi: Cuba, southeastern coast in Granma and Santiago de Cuba provinces, and probably on all the northern slopes of the Sierra Maestra. Type-locality: Ocujal, Santiago de Cuba Province, Cuba.

kingi: Bahama Is.: Andros I., Gibson Cay, Big Wood Cay; intergrades with A. a. vulturnus on Mangrove Cay. Type-locality: Gibson Cay, mouth of South Bight, Andros Island, Bahama Islands.

llanensis: Cuba, serpentine savannas in Camagüey Province. Type-locality: Just south of west end of Sierra de Cubitas, Camagiiey Province, Cuba.

marcida: Isla de la Juventud, south of the Ciénaga de Lanier. Type-locality: Jacksonville, Isla de la Juventud, Cuba.

multilineata: Bahama Is.: Berry Is. Type-locality: Bond's Cay, Berry Islands, Bahama Islands.

nigriventris: Isla de la Juventud, NE of the Ciénaga de Lanier.
Type locality: Outskirts of Cayo Piedras, Isla de la Juventud, Cuba.

obsoleta: Bahama Is.: Long I., Exuma Cays. Type-locality: Adderly's, Long Island, Bahama Islands.

orlandoi: Cuba, Archipiélago de Sabana-Camagüey; also at Buena Vista, Ciego de Avila Province. Type-locality: Cayo Santa Maria, Archipielago de Sabana-Camagliey, Villa Clara Province, Cuba.

parvinsulae: Bahama Is.: Green Cay. Type-locality: Green Cay, Bahama Islands.

paulsoni: Cuba, southern Pinar del Rio Province. Type-locality: 1 km N Las Canas, Pinar del Rio Province, Cuba.

peradusta: Cuba, from Juragmá to Calimete, thence to about 25 km B Playa Girón, Peninsula de Zapata, and inland to northern Cienfuegos Province. Type-locality: Juragua, Cienfuegos Province, Cuba.

procer: Cuba, throughout much of Pinar del Rio Province. Type-locality: San Vicente, Pinar del Rio Province, Cuba.

pullata. Cuba, northern Matanzas Province. Type-locality: 13 km NE Matanzas, Matanzas Province, Cuba.

richmondi: Bahama Is.: Bimini Is. Type-locality: Near Lyons, North Bimini Island, Bahama Islands.

sabulicolor: Cuba, southeastern coast. Type-locality: 2.8 mi. E Imias, Guantanamo Province, Cuba.

sanfelipensis: Cuba, Cayos de San Felipe. Type-locality: Cayo
Real, Cayos de San Felipe, Pinar del Rio Province, Cuba.

schwartzi: Cuba, northeastern Pinar del Rio Province. Type-locality: Henequenera de Mariel, Zona Mosquito, Pinar del Rio Province, Cuba.

secta: Isla de la Juventud, north of the Ciénaga de Lanier, except range of A. a. nigriventris. Type locality: Playa de Rocas, between Bibijagua and Ju.caro, Isla de la Juven- tud, Cuba.

sideroxylon: Bahama Is.: Andros I. Type-locality: 4.8 km N Mastic Point, Andros Island, Bahama Islands.

sublesta: Cuba, Archipiélago de Sabana-Camagüey. Type-locality: Playa Bonita, east end, Cayo Sabinal, Camagiiey Province, Cuba.

thoracica: Bahama Is.: New Providence I., Rose I., Eleuthera I., Eleuthera Cays, Little San Salvador I. Type locality: New Providence Island, Bahama Islands.

ustulata: Cuba, southeastern coast between Bahia de Santiago and Bahia de GuantAnamo, and inland north of Santiago de Cuba. Type-locality: 7.8 mi. E Siboney, Santiago de Cuba Province, Cuba.

vulturnus: Bahama Is.: Andros I. Type-locality: Marsh Bay, Andros Island, Bahama Islands.

zugi: Cuba, Archipiélago de los Canarreos, and Peninsula de Zapata. Type-locality: Cayo Largo del Sur, Archipielago de los Canarreos, Isla de la Juventud, Cuba.

INCERTAE SEDIS: Material from Herradura, Pinar del Rio Province; Cayo Macho de Tierra, about 10 mi. SE Casilda; Puerto Padre and Gibara, Holguin Province; Sierra de Jatibonico, Sabana San Felipe, Arroyo Blanco, Isla de Turiguanó is unassigned subspecifically.  
Reproductionoviparous 
TypesSyntypes: MNHN 1112, MNHN 2647, MNHN 1788, MNHN 4178 (see Schwartz, 1970a) (auberi)
Holotype: ANSP 9158 [thoracica]
Holotype: AMNH 96331 [abducta]
Holotype: AMNH 78035 [atrothorax]
Holotype: CM 60540 [behringensis]
Holotype: CM 40985 [bilateralis]
Holotype: AMNH 83028 [cacuminis]
Holotype: AMNH 96375 [itra]
Holotype: AMNH 79202 [denticola]
Holotype: IZ 52 [extorris]
Holotype: AMNH 82982 [extraria]
Holotype: CM 20440 [auberifelis]
Holotype: CM 41147 [auberifocalis]
Holotype: AMNH 78058 [galbiceps]
Holotype: IZ 96 [garridoi ]
Holotype: AMNH 82972 [gemmea]
Holotype: AMNH 83784 [granti]
Holotype: USNM 138468 [hardyi ]
Holotype: CM 40915 [kingi]
lHolotype: MCZ 59321 [lanensis]
Holotype: AMNH 82991 [marcida]
Holotype: CM 41196 [multilineata]
Holotype: IZ 5778 [nigriventris]
Holotype: CM 43976 [obsoleta]
Holotype: IZ 3427 [orlandoi]
Holotype: CM 60542 [parvinsulae]
Holotype: AMNH 83012 [paulsoni]
Holotype: IZ 85 [peradusta]
Holotype: AMNH 78390 [procer]
Holotype: AMNH 82953 [pullata]
Holotype: CM 34140 [richmnondi]
Holotype: AMNH 83941 [sabulicolor]
Holotype: IZ 2987 [sanfelipensis]
Holotype: IZ 4389 [schwartzi]
Holotype: AMNH 82997 [secta]
Holotype: CM 60539 [sideroxylon]
Holotype: AMNIR 96393 [sublesta]
Holotype: ANSP 9158 [thoracica]
Holotype: AMNH 83778 [ustulata]
Holotype: CM 60541 [vulturnus]
Holotype: AMNH 82003 [zugi] 
CommentIllustrations: Barbour and Ramsden, 1919; Schwartz and Henderson, 1985.

Subspecies: RODRIGUEZ-SCHETTINO et al. 2013 do not distinguish between subspecies, but see Schwartz 1970: 113 for a map of the Cuban subspecies. 
References
  • Barbour, T. 1904. Batrachia and Reptilia from the Bahamas. Bull. Mus. Comp. Zool. Harvard 46: 55—61 - get paper here
  • Barbour, T. and B. Shreve. 1935. Concerning some Bahamian reptiles, with notes on the fauna. Proc. Boston Soc. Nat. Hist. 40: 347-365.
  • Barbour, T. and C. T. Ramsden. 1919. The herpetology of Cuba. Mem. Mus. Comp. Zool., Harvard Coll. 47: 69-213. - get paper here
  • Cocteau, J.-T. and G. Bibron 1838. Reptiles, in R. de la Sagra. Historia Física, Política y Natural de la Isla de Cuba, Tomo IV. Arthus Bertrand, Paris, 142 pp. [1838-1843 - index on page 143] - get paper here
  • Cope, E.D. 1863. Synopsis of the species of Holcosus and Ameiva, with diagnoses of new West Indian and South American Colubridae. Proc. Acad. Nat. Sci. Philadelphia 14 [1862]: 60-82. - get paper here
  • Duméril, A. M. C. and G. Bibron. 1839. Erpétologie Générale on Histoire Naturelle Complète des Reptiles. Vol.5. Roret/Fain et Thunot, Paris, 871 pp. - get paper here
  • Estrada, A.R. 1998. Anfibios y Reptiles Encontrados Durante 1988 y 1989 en Cayo Paredón Grande, Archipiélago de Sabana-Camagüey, Cuba. Carib. J. Sci. 34 (1-2): 106-112
  • Franz, Richard;Dodd, C. Kenneth, Jr 1994. Ameiva auberi obsoleta (Exuma Islands ameiva). Bahamas: Great Bahama Bank. Herpetological Review 25 (4): 163-164 - get paper here
  • Gali and Garrido 1987. Two new subspecies of Ameiva auberi (Reptilia: Teiidae) from Cuba. Carib J. Sci. 22 (3-4):169
  • Garrido, O. H. 1975. Nuevos reptiles del archipielago cubano. Poeyana 141:1-58.
  • Goicoechea, N., Frost, D. R., De la Riva, I., Pellegrino, K. C. M., Sites, J., Rodrigues, M. T. and Padial, J. M. 2016. Molecular systematics of teioid lizards (Teioidea/Gymnophthalmoidea: Squamata) based on the analysis of 48 loci under tree-alignment and similarity-alignment. Cladistics, doi: 10.1111/cla.12150 - get paper here
  • Hower, Lindsey M., and S. Blair Hedges 2003. Molecular phylogeny and biogeography of West Indian Teiid lizards of the genus Ameiva. Carib. J. Sci. 39 (3):298-306. - get paper here
  • Lee, Daniel K.;Schwartz, Albert 1985. Four new subspecies of Ameiva auberi (Sauria, Teiidae) from the Bahama Islands. Annals of the Carnegie Museum 54 (2): 11-21
  • Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96 - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Schwartz, A. & McCoy, C.J. 1970. A systematic review of Ameiva auberi Cocteau (Reptilia, Teiidae) in Cuba and the Bahamas. I. The Cuban subspecies. II: The Bahamian subspecies. III. Discussion. Annals of the Carnegie Museum 41(4): 45-168.
  • Schwartz, A. and Henderson, R.W. 1985. A guide to the identification of the amphibians and reptiles of the West Indies exclusive of Hispaniola. Milwaukee Public Mus., 165 pp.
  • Schwartz, A. and McCoy, C.J. 1975. A New Name for a Cuban Ameiva (Reptilia: Teiidae) Herpetologica 31(2): 240 - get paper here
  • Tucker, Derek B.; | Stephen Blair Hedges | Guarino R. Colli | Robert Alexander Pyron | Jack W. Sites Jr 2017. Genomic timetree and historical biogeography of Caribbean island ameiva lizards (Pholidoscelis: Teiidae) Ecology and Evolution 7:7080–7090.
  • Varona, Luis S.;Garrido, Orlando H. 1970. Vertebrados de los Cayos de San Felipe, Cuba, Inclyendo una nueva especie de Jutía. Poeyana (75): 1-26
 
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