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Phrynosoma modestum GIRARD, 1852

IUCN Red List - Phrynosoma modestum - Least Concern, LC

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Higher TaxaPhrynosomatidae, Phrynosomatinae, Phrynosomatini; Iguania, Sauria, Squamata (lizards) 
Common NamesE: Roundtail Horned Lizard
G: Rundschwanz-Krötenechse 
SynonymPhrynosoma modestum GIRARD in BAIRD & GIRARD 1852: 69
Doliosaurus modestus — GIRARD 1858: 409
Phrynosoma modestum — GENTRY 1885: 148
Phrynosoma modestum — BOULENGER 1885: 248
Anota modesta — COPE 1896: 1014
Phrynosoma modestum — SMITH & TAYLOR 1950: 101
Phrynosoma modestum — STEBBINS 1985: 142
Phrynosoma modestum — CONANT & COLLINS 1991: 114
Phrynosoma modestum — LINER 1994
Phrynosoma (Doliosaurus) modestum — LINER 2007
Phrynosoma (Doliosaurus) modestum — DE QUEIROZ in CROTHER et al. 2017 
DistributionUSA (SE Arizona, New Mexico, W Texas),
Mexico (Chihuahua, Coahuila, W Nuevo Leon, NE Durango, NE Zacatecas, NW San Luis Potosi, Sonora, Aguascalientes; one isolated record in W Tamaulipas)

Type locality: The Rio Grande west of San Antonio, Texas [CHURCHILL], from between San Antonio and El Paso [GRAHAM]. Restricted to Las Cruces, New Mexico by SMITH & TAYLOR 1950.  
TypesSyntype. UIMNH 40746; J. H. Clark; 1857; maybe USNM (fide SMITH & TAYLOR 1950) 
CommentGroup: Belongs to the Doliosaurus clade fide LEACHE & MCGUIRE 2006. 
  • Baird,S.F. and Girard,C. 1852. Characteristics of some new reptiles in the Museum of the Smithsonian Institution. Proc. Acad. Nat. Sci. Philadelphia 6: 68-70 - get paper here
  • Baird,S.F. and Girard,C. 1852. Reptiles. in: Stansbury’s Exploration and survey of the valley of the Great Salt Lake of Utah, 1852, including a Reconnoissance of a new Route through the Rocky Mountains. pp. 336-365
  • Bartlett, R. D. & Bartlett, P. 1999. A Field Guide to Texas Reptiles and Amphibians. Gulf Publishing Co., Houston, Texas, 331 pp.
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Burt, Charles E. 1935. Further records of the ecology and distribution of amphibians and reptiles in the middle west. American Midland Naturalist 16 (3): 311-336 - get paper here
  • Conant,R. & Collins,J.T. 1991. A Field Guide to Reptiles and Amphibians of Eastern/Central North America, 3rd ed. Houghton Mifflin (Boston/New York), xx + 450 p.
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
  • Davis DR, LaDuc TJ 2018. Amphibians and reptiles of C. E. Miller Ranch and the Sierra Vieja, Chihuahuan Desert, Texas, USA ZooKeys 735: 97-130 - get paper here
  • Degenhardt, William G.; C. W. Painter, and A. H. Price 1996. Amphibians and reptiles of New Mexico. Univ. New Mexico Press, 431 pp.
  • Dixon, James R. 2000. Amphibians and reptiles of Texas, second edition. Texas A&M University Press, 421 pp.
  • Fabian, B. F 2014. Überlegungen zur Ernährung von Krötenechsen. Reptilia (Münster) 19 (107): 24-29
  • García-De la Peña, Cristina; Héctor Gadsden, Rosalinda Palomo-Ramos, Ana B. Gatica-Colima, Pablo A. Lavín-Murcio, and Gamaliel Castañeda. 2012. Spatial Segregation of Microhabitats Within a Community of Lizards in Médanos de Samalayuca, Chihuahua, Mexico. Southwestern Naturalist 57 (4): 430-434 - get paper here
  • Gentry, A.F. 1885. A review of the genus Phrynosoma. Proc. Acad. Nat. Sci. Philad. (ser. 3) 37: 138-148 - get paper here
  • Jones, L.L. & Lovich, R.E. 2009. Lizards of the American Southwest. A photographic field guide. Rio Nuevo Publishers, Tucson, AZ, 568 pp. [review in Reptilia 86: 84] - get paper here
  • Kasper, Stephen. 2014. Geographical Distribution: Phrynosoma modestum (round-tailed horned lizard). Herpetological Review 45 (4): 660
  • Lara-Reséndiz Rafael A., Arenas-Moreno Diego M., Beltrán-Sánchez Elizabeth, Gramajo Weendii, Verdugo-Molina Javier, Sherbrooke Wade C. et al. 2015. Selected body temperature of nine species of Mexican horned lizards (Phrynosoma) Rev. Mex. Biodiv. 86 (1): 275-278. - get paper here
  • Leaché, Adam D. and Jimmy A. McGuire 2006. Phylogenetic relationships of horned lizards (Phrynosoma ) based on nuclear and mitochondrial data: Evidence for a misleading mitochondrial gene tree. Molecular Phylogenetics and Evolution 39 (3): 628-644 - get paper here
  • Lemos-Espinal, J.A. & Smith, H.M. 2007. Amphibians and reptiles of the state of Chihuahua, Mexico. Universidad Nacional Autonoma de Mexico, 613 pp.
  • Lemos-Espinal, J.A. & Smith, H.M. 2007. Amphibians and reptiles of the state of Coahuila, Mexico. Universidad Nacional Autonoma de Mexico, 550 pp.
  • Lemos-Espinal, Julio A. and James R. Dixon 2013. Amphibians and Reptiles of San Luis Potosí. Eagle Mountain Publishing, xii + 300 pp.
  • Lemos-Espinal, Julio A., David Chiszar, and Hobart M. Smith 2004. Selected Records of 2003 Lizards from Chihuahua and Sonora, Mexico. Bull. Chicago Herp. Soc. 39 (9): 164-168 - get paper here
  • Liner, Ernest A. 2007. A CHECKLIST OF THE AMPHIBIANS AND REPTILES OF MEXICO. Louisiana State University Occasional Papers of the Museum of Natural Science 80: 1-60 - get paper here
  • Liner, Ernest A.;Montanucci, Richard R.;González-Alonso, Arturo;Mendoza Quijano, Fernando 1993. An additional contribution to the herpetology of Northern Coahuila, Mexico. Boletín de la Sociedad Herpetológica Mexicana 5 (1): 9-11 - get paper here
  • Mayne, Peter J. 1997. Geographic Distribution. Phrynosoma modestum. Herpetological Review 28 (1): 50 - get paper here
  • McCranie, J.R. & Wilson, L.D. 2001. The herpetofauna of the Mexican State of Aguascalientes. Courier Forschungsinstitut Senckenberg 230: 1-57
  • Meyers, J.J.; Herrel, A. & Nishikawa, K. 2006. Morphological correlates of ant eating in horned lizards (Phrynosoma). Biological Journal of the Linnean Society 89: 13–24 - get paper here
  • Quintero-Díaz, Gustavo E., Armando Cardona-Arceo and Rubén Alonso Carbajal-Márquez. 2015. Phrynosoma modestum Girard, 1852. Mexico, Aguascalientes. Mesoamerican Herpetology 2(3): 355 - get paper here
  • Reeder,T.W. & Montanucci,R.R. 2001. Phylogenetic analysis of the horned lizards (Phrynomomatidae: Phrynosoma): evidence from mitochondrial DNA and morphology. Copeia 2001 (2): 309-323 - get paper here
  • Sherbrooke, Wade C. 2003. Introduction to Horned Lizards of North America. University of California Press, Berkeley, 178 pp.
  • Sherbrooke, Wade C. 2017. Antipredator Nest Guarding by Female Horned Lizards (Phrynosoma): Iguanian Parental Care Herpetologica Dec 2017, Vol. 73, No. 4: 331-337. - get paper here
  • Sherbrooke, Wade C. Frost, Sally Kay Viparina. 1989. Integumental chromatophores of a color-change, thermoregulating lizard, Phrynosoma modestum (Iguanidae, Reptilia). American Museum Novitates (2943): 1-14 - get paper here
  • Sherbrooke, Wade C.;Montanucci, Richard R. 1988. Stone mimicry in the round-tailed horned lizard, Phrynosoma modestum (Sauria: Iguanidae). Journal of Arid Environments 14: 275-284
  • Smith, H.M. 1935. Miscellaneous notes on Mexican lizards. Univ. Kansas Sci. Bull. 22: 119-156 - get paper here
  • Smith, Hobart M. 1934. Notes on some lizards of the genus Phrynosoma from Mexico. Transactions of the Kansas Academy of Science 37: 287-297 - get paper here
  • Stebbins,R.C. 1985. A Field Guide to Western Reptiles and Amphibians, 2nd ed. Houghton Mifflin, Boston
  • Tanner, Wilmer W. 1958. Herpetological range extensions. Herpetologica 14: 195-196 - get paper here
  • Terán-Juárez, Sergio A., Elí García Padilla, Vicente Mata-Silva, Jerry D. Johnson and Larry David Wilson. 2016. The herpetofauna of Tamaulipas, Mexico: composition, distribution, and conservation status. Mesoamerican Herpetology 3 (1): 43–113 - get paper here
  • Valdez-Lares, R.; R. Muñiz-Martínez; E.Gadsden; G. Aguirre-León; G. Castañeda-Gaytán; R. Gonzalez-Trápaga 2013. Checklist of amphibians and reptiles of the state of Durango, México. Check List 9 (4):714-724 - get paper here
  • Vázquez Díaz, Joel;Quintero Díaz, Gustavo E. 2005. Anfibios y Reptiles de Aguascalientes [2nd ed.]. CONABIO, CIEMA, 318 pp.
  • Wernicke, Jörg & Klaus Haase 2014. Phrynosoma modestum. Kleine Zwerge mit interessantem Verhalten. Terraria-Elaphe 2014 (2): 34-37 - get paper here
  • Werning, H. 2014. Krötenechsen – eine (sehr gute) Laune der Natur. Reptilia (Münster) 19 (107): 16-23
  • Werning, H. 2015. Jurassic World im Wohnzimmer. Mini-Dinos für das Terrarium Reptilia (Münster) 20 (114): 26-37 - get paper here
  • Werning, H. 2018. Nordamerikanische Kleinleguane – Die Familie Phrynosomatidae Reptilia 23 (131): 14-19
  • Werning, Heiko 2012. Die Reptilien und Amphibien des Südwestens. Draco 13 (50): 18-60 - get paper here
  • Whiting M J. Dixon J R. 1996. Phrynosoma modestum. Catalogue of American Amphibians and Reptiles (630): 1-6. - get paper here
  • Williams, Kenneth L.;Smith, Hobart M.;Chrapliwy, Pete S. 1960. Turtles and lizards from Northern Mexico. Transactions of the Illinois State Academy of Science 53 (1/2): 36-45
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