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Phrynosoma modestum GIRARD, 1852

IUCN Red List - Phrynosoma modestum - Least Concern, LC

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Higher TaxaPhrynosomatidae, Phrynosomatinae, Phrynosomatini; Iguania, Sauria, Squamata (lizards) 
Subspecies 
Common NamesE: Roundtail Horned Lizard
G: Rundschwanz-Krötenechse 
SynonymPhrynosoma modestum GIRARD in BAIRD & GIRARD 1852: 69
Doliosaurus modestus — GIRARD 1858: 409
Phrynosoma modestum — GENTRY 1885: 148
Phrynosoma modestum — BOULENGER 1885: 248
Anota modesta — COPE 1896: 1014
Phrynosoma modestum — SMITH & TAYLOR 1950: 101
Phrynosoma modestum — STEBBINS 1985: 142
Phrynosoma modestum — CONANT & COLLINS 1991: 114
Phrynosoma modestum — LINER 1994
Phrynosoma (Doliosaurus) modestum — LINER 2007
Phrynosoma (Doliosaurus) modestum — DE QUEIROZ in CROTHER et al. 2017 
DistributionUSA (SE Arizona, New Mexico, W Texas),
Mexico (Chihuahua, Coahuila, W Nuevo Leon, NE Durango, NE Zacatecas, NW San Luis Potosi, Sonora, Aguascalientes; one isolated record in W Tamaulipas)

Type locality: The Rio Grande west of San Antonio, Texas [CHURCHILL], from between San Antonio and El Paso [GRAHAM]. Restricted to Las Cruces, New Mexico by SMITH & TAYLOR 1950.  
Reproductionoviparous. 
TypesSyntypes: INHS (= UIMNH) 40746; J. H. Clark; 1857; USNM 164 (see also SMITH & TAYLOR 1950); USNM 165660 was intended to be the lectotype. Lectotype designation by Ralph Axtell but designation is not valid as it was never published (pers. comm., Ralph Axtell, July 1988, cited in USNM catalog). 
Diagnosis 
CommentGroup: Belongs to the Doliosaurus clade fide LEACHE & MCGUIRE 2006. 
Etymology 
References
  • Aguilar-Morales, Cecilia and Thomas R. Van Devender 2018. Horned Lizards (Phrynosoma) of Sonora, Mexico: Distribution and Ecology. Sonoran Herpetologist 31 (3): 40-50 - get paper here
  • Axtell, Ralph W. 1939. AMPHIBIANS AND REPTILES OF THE BLACK GAP WILDLIFE MANAGEMENT AREA, BREWSTER COUNTY, TEXAS. Southwestern Naturalist 4 (2): 88-109
  • Baird,S.F. and Girard,C. 1852. Reptiles. in: Stansbury’s Exploration and survey of the valley of the Great Salt Lake of Utah, 1852, including a Reconnoissance of a new Route through the Rocky Mountains. pp. 336-365
  • Baird,S.F. and Girard,C. 1852. Characteristics of some new reptiles in the Museum of the Smithsonian Institution. Proc. Acad. Nat. Sci. Philadelphia 6: 68-70 - get paper here
  • Bartlett, R. D. & Bartlett, P. 1999. A Field Guide to Texas Reptiles and Amphibians. Gulf Publishing Co., Houston, Texas, 331 pp.
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Burt, Charles E. 1935. Further records of the ecology and distribution of amphibians and reptiles in the middle west. American Midland Naturalist 16 (3): 311-336 - get paper here
  • Conant,R. & Collins,J.T. 1991. A Field Guide to Reptiles and Amphibians of Eastern/Central North America, 3rd ed. Houghton Mifflin (Boston/New York), xx + 450 p.
  • Cooper Jr, William E. and Wade C. Sherbrooke 2018. Crypsis and Flight Initiation Distance in Horned Lizards Sonoran Herpetologist 31 (3): 51-55 - get paper here
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
  • Davis DR, LaDuc TJ 2018. Amphibians and reptiles of C. E. Miller Ranch and the Sierra Vieja, Chihuahuan Desert, Texas, USA ZooKeys 735: 97-130 - get paper here
  • Degenhardt, William G.; C. W. Painter, and A. H. Price 1996. Amphibians and reptiles of New Mexico. Univ. New Mexico Press, 431 pp.
  • Dixon, James R. 2000. Amphibians and reptiles of Texas, second edition. Texas A&M University Press, 421 pp.
  • Fabian, B. F 2014. Überlegungen zur Ernährung von Krötenechsen. Reptilia (Münster) 19 (107): 24-29
  • García-De la Peña, Cristina; Héctor Gadsden, Rosalinda Palomo-Ramos, Ana B. Gatica-Colima, Pablo A. Lavín-Murcio, and Gamaliel Castañeda. 2012. Spatial Segregation of Microhabitats Within a Community of Lizards in Médanos de Samalayuca, Chihuahua, Mexico. Southwestern Naturalist 57 (4): 430-434 - get paper here
  • Gentry, A.F. 1885. A review of the genus Phrynosoma. Proc. Acad. Nat. Sci. Philad. (ser. 3) 37: 138-148 - get paper here
  • Hernandez T, Herr MW, Stevens S, Cork K, Medina-Nava C, Vialpando CJ, Warfel T, Fields N, Brodie C, Graham SP 2019. New distribution records for amphibians and reptiles in eastern Chihuahua, Mexico. Check List 15(1): 79-86 - get paper here
  • Jones, L.L. & Lovich, R.E. 2009. Lizards of the American Southwest. A photographic field guide. Rio Nuevo Publishers, Tucson, AZ, 568 pp. [review in Reptilia 86: 84] - get paper here
  • Kasper, Stephen. 2014. Geographical Distribution: Phrynosoma modestum (round-tailed horned lizard). Herpetological Review 45 (4): 660 - get paper here
  • Lara-Reséndiz Rafael A., Arenas-Moreno Diego M., Beltrán-Sánchez Elizabeth, Gramajo Weendii, Verdugo-Molina Javier, Sherbrooke Wade C. et al. 2015. Selected body temperature of nine species of Mexican horned lizards (Phrynosoma) Rev. Mex. Biodiv. 86 (1): 275-278. - get paper here
  • Leaché, Adam D. and Jimmy A. McGuire 2006. Phylogenetic relationships of horned lizards (Phrynosoma ) based on nuclear and mitochondrial data: Evidence for a misleading mitochondrial gene tree. Molecular Phylogenetics and Evolution 39 (3): 628-644 - get paper here
  • Lemos-Espinal, J.A. & Smith, H.M. 2007. Amphibians and reptiles of the state of Chihuahua, Mexico. Universidad Nacional Autonoma de Mexico, 613 pp.
  • Lemos-Espinal, J.A. & Smith, H.M. 2007. Amphibians and reptiles of the state of Coahuila, Mexico. Universidad Nacional Autonoma de Mexico, 550 pp.
  • Lemos-Espinal, Julio A. and James R. Dixon 2013. Amphibians and Reptiles of San Luis Potosí. Eagle Mountain Publishing, xii + 300 pp.
  • Lemos-Espinal, Julio A., David Chiszar, and Hobart M. Smith 2004. Selected Records of 2003 Lizards from Chihuahua and Sonora, Mexico. Bull. Chicago Herp. Soc. 39 (9): 164-168 - get paper here
  • Liner, Ernest A. 2007. A CHECKLIST OF THE AMPHIBIANS AND REPTILES OF MEXICO. Louisiana State University Occasional Papers of the Museum of Natural Science 80: 1-60 - get paper here
  • Liner, Ernest A.;Montanucci, Richard R.;González-Alonso, Arturo;Mendoza Quijano, Fernando 1993. An additional contribution to the herpetology of Northern Coahuila, Mexico. Boletín de la Sociedad Herpetológica Mexicana 5 (1): 9-11 - get paper here
  • Mayne, Peter J. 1997. Geographic Distribution. Phrynosoma modestum. Herpetological Review 28 (1): 50 - get paper here
  • McCranie, J.R. & Wilson, L.D. 2001. The herpetofauna of the Mexican State of Aguascalientes. Courier Forschungsinstitut Senckenberg 230: 1-57
  • Meyers, J.J.; Herrel, A. & Nishikawa, K. 2006. Morphological correlates of ant eating in horned lizards (Phrynosoma). Biological Journal of the Linnean Society 89: 13–24 - get paper here
  • Nevárez-de-los-Reyes, Manuel, David Lazcano, Elí García-Padilla, Vicente Mata-Silva, Jerry D. Johnson and Larry David Wilson. 2016. The Herpetofauna of Nuevo León, Mexico: Composition, Distribution, and Conservation. Mesoamerican Herpetology 3 (3): 558–638 - get paper here
  • Quintero-Díaz, Gustavo E., Armando Cardona-Arceo and Rubén Alonso Carbajal-Márquez. 2015. Phrynosoma modestum Girard, 1852. Mexico, Aguascalientes. Mesoamerican Herpetology 2(3): 355 - get paper here
  • Reeder,T.W. & Montanucci,R.R. 2001. Phylogenetic analysis of the horned lizards (Phrynomomatidae: Phrynosoma): evidence from mitochondrial DNA and morphology. Copeia 2001 (2): 309-323 - get paper here
  • Sherbrooke, Wade C. 2003. Introduction to Horned Lizards of North America. University of California Press, Berkeley, 178 pp.
  • Sherbrooke, Wade C. 2017. Antipredator Nest Guarding by Female Horned Lizards (Phrynosoma): Iguanian Parental Care Herpetologica Dec 2017, Vol. 73, No. 4: 331-337. - get paper here
  • Sherbrooke, Wade C. Frost, Sally Kay Viparina. 1989. Integumental chromatophores of a color-change, thermoregulating lizard, Phrynosoma modestum (Iguanidae, Reptilia). American Museum Novitates (2943): 1-14 - get paper here
  • Sherbrooke, Wade C.;Montanucci, Richard R. 1988. Stone mimicry in the round-tailed horned lizard, Phrynosoma modestum (Sauria: Iguanidae). Journal of Arid Environments 14: 275-284
  • Smith, H.M. 1935. Miscellaneous notes on Mexican lizards. Univ. Kansas Sci. Bull. 22: 119-156 - get paper here
  • Smith, Hobart M. 1934. Notes on some lizards of the genus Phrynosoma from Mexico. Transactions of the Kansas Academy of Science 37: 287-297 - get paper here
  • Stebbins,R.C. 1985. A Field Guide to Western Reptiles and Amphibians, 2nd ed. Houghton Mifflin, Boston
  • Tanner, Wilmer W. 1958. Herpetological range extensions. Herpetologica 14: 195-196 - get paper here
  • Terán-Juárez, Sergio A., Elí García Padilla, Vicente Mata-Silva, Jerry D. Johnson and Larry David Wilson. 2016. The herpetofauna of Tamaulipas, Mexico: composition, distribution, and conservation status. Mesoamerican Herpetology 3 (1): 43–113 - get paper here
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  • Vázquez Díaz, Joel;Quintero Díaz, Gustavo E. 2005. Anfibios y Reptiles de Aguascalientes [2nd ed.]. CONABIO, CIEMA, 318 pp.
  • Wernicke, Jörg & Klaus Haase 2014. Phrynosoma modestum. Kleine Zwerge mit interessantem Verhalten. Terraria-Elaphe 2014 (2): 34-37 - get paper here
  • Werning, H. 2014. Krötenechsen – eine (sehr gute) Laune der Natur. Reptilia (Münster) 19 (107): 16-23
  • Werning, H. 2015. Jurassic World im Wohnzimmer. Mini-Dinos für das Terrarium Reptilia (Münster) 20 (114): 26-37 - get paper here
  • Werning, H. 2018. Nordamerikanische Kleinleguane – Die Familie Phrynosomatidae Reptilia 23 (131): 14-19
  • Werning, Heiko 2012. Die Reptilien und Amphibien des Südwestens. Draco 13 (50): 18-60 - get paper here
  • Whiting M J. Dixon J R. 1996. Phrynosoma modestum. Catalogue of American Amphibians and Reptiles (630): 1-6. - get paper here
  • Williams, Kenneth L.;Smith, Hobart M.;Chrapliwy, Pete S. 1960. Turtles and lizards from Northern Mexico. Transactions of the Illinois State Academy of Science 53 (1/2): 36-45
 
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