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Higher TaxaPhyllodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Simpson’s Leaf-toed Gecko, Western Galápagos Leaf-toed Gecko
S: Geco de Simpson, salamanquesa de Simpson, geco occidental de Galápagos. 
SynonymPhyllodactylus simpsoni ARTEAGA, BUSTAMANTE, VIEIRA, TAPIA & GUAYASAMIN 2019 
DistributionEcuador: Galapagos Island (Isabela Island, Fernandina Island)

Type locality: Puerto Villamil (-0.95622, -90.97023; 3 m), Isabela Island, Galápagos, Ecuador  
TypesHolotype. CAS 10339, an adult of undetermined sex collected by Joseph Slevin, on November 1, 1905.
Paratypes. CAS 11245, an adult of undetermined sex collected by Joseph Slevin at the type locality on March 9, 1906. CAS 10351, an adult of undetermined sex collected by Francis Williams at the type locality on November 3, 1905. 
DiagnosisDiagnosis: males 10 cm, females 9.6 cm. Phyllodactylus simpsoni is placed in the genus Phyllodactylus based on phylogenetic evidence (Fig. 1). The species is compared to other geckos traditionally assigned to P. galapagensis. From P. galapagensis, it differs in lacking pointed tubercles on the top of the head and having fewer (71–80 vs 83–103) midbody scales (see Lanza [1973]2 for an explanation of this measurement). From P. maresi and P. duncanensis, it differs in having pine-cone-shaped and symmetrical fingertips, as opposed to asymmetrical pine-cone-shaped fingertips (Fig. 6). From P. andysabini, it differs in having supranasals occasionally (in 8 out of 14 individuals examined by us) in contact (versus in not in contact in 9 out of 9 P. andysabini; Fig. 7), and having the throat usually (in 9 out of 14 individuals) immaculate as opposed to densely stippled with dark brown pigment (Fig. 8). Genetic divergence in a 181 bp long fragment of the mitochondrial 12S gene between the sister species P. simpsoni and P. andysabini is 3–5%, whereas intraspecific distances are less than 2.3% in six individuals of P. simpsoni (Arteaga et al. 2019: 184). 
EtymologyThe specific name simpsoni honors Dr. Nigel Simpson for his long-standing and visionary leadership in conservation. Nigel is a founding board member of the Ecuadorian conservation organizations Fundación Jocotoco and Fundación Ecominga. 
  • Arteaga A, Bustamante L, Vieira J, Tapia W, Guayasamin JM (Eds) 2019. Reptiles of Galápagos. Tropical Herping - get paper here
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