Podarcis carbonelli PÉREZ-MELLADO, 1981
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|Higher Taxa||Lacertidae, Lacertinae, Sauria, Lacertoidea, Squamata (lizards)|
|Subspecies||Podarcis carbonelli berlengensis VICENTE 1985|
Podarcis carbonelli carbonelli PEREZ MELLADO 1981
|Common Names||E: Carbonell’s Wall Lizard|
|Synonym||Podarcis bocagei carbonelli PÉREZ MELLADO 1981|
Podarcis bocagei carbonelli — SÁ-SOUSA 1999
Podarcis carbonelli — SÁ-SOUSA & HARRIS 2002
Podarcis carbonelli — HARRIS et al. 2002
Podarcis carbonelli — KALIONTZOPOULOU et al. 2007
Podarcis carbonelli — LIVIGNI 2013: 290
Podarcis carbonellae — BEOLENS et al. 2011: 47
Podarcis carbonelli — SPEYBROECK et al. 2020
Podarcis bocagei berlengensis VICENTE, 1985
Podarcis bocagei berlengensis — MALKMUS 1995
Podarcis carbonelli berlengensis — HARRIS & SÁ-SOUSA 2001
Podarcis carbonelli berlengensis — SINDACO & JEREMČENKO 2008
|Distribution||Spain (dunes of Coto Doñana, south of Andalusia,|
more than 400 km south of the Sistema Central range (Magraner, 1986); this was often refuted (Peréz-Mellado, 1997, 1998; Barbadillo et al., 1999); mountains to the
Sistema Central and along the Atlantic lowlands (Sá-Sousa 1999, 2000).
berlengensis: Berlengas islands off the western coast of Portugal (Vicente, 1985); Type locality: Berlenga Island, Portugal.
Type locality: Laguna de San Marcos, La Alberca, Prov. Salamanca, Spain.
|Types||Holotype: MNCN (Madrid), fide V. Pérez-Mellado, pers. comm., 16 Apr 2019; given in original description as VPM 07010680, adult male (= private collection Valentin Pérez-Mellado).|
Holotype: MNHN-RA 1982.0995 (by original desingation; paratypes: MNHN-RA 1982.0996-1024, MNHn-RA 1982.1025-1039 and MNHN-RA 1983.0001-0005 [berlengensis]
|Diagnosis||Diagnosis. According to Pérez-Mellado (1981) P. carbonelli is a small lizard with SVL <63.5 mm, less robust than P. bocagei. Head (pileous) is relatively short. The mid-back coloration is often uniform brown, sometimes with a series of small black points in the vertebral zone. The light dorsal stripes are bright green with irregular edges in case of the males while having regular ones in the females. The mid-back coloration during the mating season shows shifts in relative frequencies from the western part towards the eastern part of the distribution range of P. carbonelli: the bright green or brown-grenish tones in the mid-back of males are more frequent in Portugal, whereas the the brown tones with green dorsal stripes predominate in the Spanish western ‘Sistema Central’. However, the concolor pattern (= single uniform brown without black pigmentation) is found in low frequence in males/females from all populations studied. Extensive geographic variation exists in body size and shape and in scalation traits, southern populations being visibly differentiated (Kaliontzopoulou et al. 2010). Some degree of local introgression with P. bocagei has been documented at the northern part of the range of P. carbonelli (Pinho et al. 2009).|
|Comment||Podarcis hispanica is a genetically (Kaliontzopoulou et al. 2011) and morphologically (Carretero 2008; Kaliontzopoulou et al. 2012) very diverse species. In fact, P. bocagei and P. (bocagei) carbonelli are nested within the P. hispanica clade based on DNA sequence data (PINHO et al. 2005; Kaliontzopoulou et al. 2011).|
Distribution: see map in Caeiro-Dias et al. 2018.
|Etymology||Named after J. Carbonell, the describer's wife. Perez-Mellado did not provide the first name of Ms. Carbonell, and actually did not use the feminine suffix. Beolens et al. 2011: 47 hence suggested to rename it to P. carbonellae.|
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