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Podarcis pityusensis (BOSCÁ, 1883)

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Higher TaxaLacertidae, Sauria, Lacertoidea, Squamata (lizards) 
SubspeciesPodarcis pityusensis affinis (MÜLLER 1927)
Podarcis pityusensis ahorcadosi (EISENTRAUT 1930)
Podarcis pityusensis calaesaladae (MÜLLER 1928)
Podarcis pityusensis canensis (EISENTRAUT 1928)
Podarcis pityusensis canaretensis CIRER 1980
Podarcis pityusensis caragolensis (BUCHHOLZ 1954)
Podarcis pityusensis carlkochi (MERTENS & MÜLLER 1940)
Podarcis pityusensis characae (BUCHHOLZ 1954)
Podarcis pityusensis formenterae (EISENTRAUT 1928)
Podarcis pityusensis frailensis (EISENTRAUT 1928)
Podarcis pityusensis gastabiensis (EISENTRAUT 1928)
Podarcis pityusensis gorrae (EISENTRAUT 1928)
Podarcis pityusensis hedwigkamerae (MÜLLER 1927)
Podarcis pityusensis hortae (BUCHHOLZ 1954)
Podarcis pityusensis kameriana (MERTENS 1927)
Podarcis pityusensis maluquerorum MERTENS 1921
Podarcis pityusensis muradae (EISENTRAUT 1928)
Podarcis pityusensis negrae (EISENTRAUT 1928)
Podarcis pityusensis pityusensis (BOSCÁ 1833)
Podarcis pityusensis puercosensis (BUCHHOLZ 1954) -- invalid
Podarcis pityusensis purroigensis (BUCHHOLZ 1954) -- invalid
Podarcis pityusensis ratae (EISENTRAUT 1928)
Podarcis pityusensis redonae (EISENTRAUT 1928)
Podarcis pityusensis sabinae (BUCHHOLZ 1954) -- invalid
Podarcis pityusensis schreitmuelleri (MÜLLER 1927)
Podarcis pityusensis subformenterae (BUCHHOLZ 1954) -- invalid
Podarcis pityusensis tagomagensis (MÜLLER 1927)
Podarcis pityusensis torretensis (BUCHHOLZ 1954)
Podarcis pityusensis vedrae (MÜLLER 1927) 
Common NamesE: Ibiza Wall Lizard
G: Pityusen-Eidechse 
SynonymLacerta muralis var. pityusensis BOSCÁ 1883: 246
Podarcis pityusensis — ENGELMANN et al 1993
Podarcis pityusensis — LIVIGNI 2013: 359

Podarcis pityusensis pityusensis (BOSCÁ 1833)
Lacerta lilfordi miguelensis EISENTRAUT 1928: 467
Podarcis pityusensis characae (BUCHHOLZ 1954)
Podarcis pityusensis pityusensis — KRONIGER & ZAWADZKI 2002

Podarcis pityusensis affinis (MÜLLER 1927)
Lacerta lilfordi affinis MÜLLER 1927: 269
Podarcis pityusensis affinis — SINDACO & JEREMČENKO 2008

Podarcis pityusensis ahorcadosi (EISENTRAUT, 1930)
Lacerta pityusensis ahorcadosi EISENTRAUT 1930: 399
Lacerta pityusensis ahorcadosi — BUCHHOLZ 1954: 80
Podarcis pityusensis ahorcadosi — BAUER & GÜNTHER 1995

Podarcis pityusensis calaesaladae (MÜLLER 1928)
Podarcis pityusensis calaesaladae — SALVADOR 1984
Podarcis pityusensis calaesaladae — SALVADOR 1986

Podarcis pityusensis canensis (EISENTRAUT, 1928)
Lacerta lilfordi canensis EISENTRAUT 1928: 466
Podarcis pityusensis canensis — SALVADOR 1984
Podarcis pityusensis canensis — SALVADOR 1986
Podarcis pityusensis canensis — BAUER & GÜNTHER 1995

Podarcis pityusensis canaretensis CIRER 1980
Podarcis pityusensis canaretensis — SALVADOR 2006

Podarcis pityusensis caragolensis (BUCHHOLZ 1954)
Lacerta pityusensis caragolensis BUCHHOLZ 1954: 81
Podarcis pityusensis caragolensis — SALVADOR 1984
Podarcis pityusensis caragolensis — SALVADOR 1986

Podarcis pityusensis carlkochi (MERTENS & MÜLLER 1940)
Lacerta liìfordi kochi MÜLLER 1927
Lacerta pityusensis carl-kochi MERTENS & MÜLLER 1940 (nomen novum)
Podarcis pityusensis carlkochi — SALVADOR 1984
Podarcis pityusensis carlkochi — SALVADOR 1986
Podarcis pityusensis carlkochi — TIEDEMANN et al. 1994

Podarcis pityusensis characae (BUCHHOLZ 1954)
Lacerta pityusensis characae BUCHHOLZ 1954: 86
Podarcis pityusensis characae — SALVADOR 1984
Podarcis pityusensis characae — SALVADOR 1986

Podarcis pityusensis formenterae (EISENTRAUT 1928)
Lacerta lilfordi formenterae EISENTRAUT 1928: 123
Lacerta lilfordi formenterae MÜLLER 1928
Lacerta lilfordi espalmadoris MÜLLER 1928
Lacerta lilfordi espardellensis EISENTRAUT 1928: 123 (fide BAUER & GÜNTHER 1995)
Lacerta lilfordi grueni MÜLLER 1928 (fide SALVADOR 1984)
Lacerta lilfordi grisea (EISENTRAUT 1928: 122; [preoccupied])
Lacerta lilfordi grueni —EISENTRAUT 1928
Lacerta pityusensis grueni — EISENTRAUT 1928
Lacerta lilfordi grueni — MERTENS 1934: 81
Lacerta lilfordi formenterae — MERTENS 1934: 83
Lacerta pityusensis formenterae — MERTENS & MÜLLER 1940: 37
Lacerta lilfordi grueni MÜLLER 1949
Lacerta pityusensis grueni — HARTMANN 1953
Lacerta pityusensis espalmadoris — BUCHHOLZ 1954
Lacerta pityusensis formenterae — BUCHHOLZ 1954
Lacerta pityusensis grueni — COLOM 1957
Lacerta pityusensis grueni — COLOM 1964
Lacerta pityusensis grueni — COMPTE SART 1966
Lacerta pityusensis grueni — RODRIGUEZ RUIZ 1976
Podarcis pityusensis grueni — MARTÍNEZ-RICA & CIRER 1982
Podarcis pityusensis formenterae — SALVADOR 1984
Lacerta lilfordi espardellensis — SALVADOR 1986: 240
Podarcis pityusensis formenterae — CIRER 1989
Podarcis lilfordi grueni — BAUER & GÜNTHER 1995
Podarcis pityusensis formenterae — PONS & PALMER 1996
Podarcis pityusensis formenterae — BAUER & GÜNTHER 1995
Podarcis pityusensis formenterae — KRONIGER & ZAWADZKI 2005
Podarcis pityusensis formenterae — SINDACO & JEREMČENKO 2008
Podarcis pityusensis formenterae — ZAWADZKI 2013
Podarcis pityusensis forerae — ZAWADZKI 2014: 23
Podarcis pityusensis formenterae — KWET & TRAPP 2014

Podarcis pityusensis frailensis (EISENTRAUT, 1928)
Lacerta lilfordi frailensis EISENTRAUT 1928: 467
Lacerta lilfordi frailensis MÜLLER 1928
Lacerta lilfordi frailensis — MERTENS 1934: 88
Podarcis pityusensis frailensis — SALVADOR 1984
Podarcis pityusensis frailensis — SALVADOR 1986
Podarcis pityusensis frailensis — BAUER & GÜNTHER 1995

Podarcis pityusensis gastabiensis (EISENTRAUT, 1928)
Lacerta lilfordi gastabiensis EISENTRAUT 1928: 123
Lacerta pityusensis gastabiensis — MERTENS & MÜLLER 1940: 37
Lacerta pityusensis gastabiensis — BUCHHOLZ 1954: 79
Podarcis pityusensis gastabiensis — SALVADOR 1984
Podarcis pityusensis gastabiensis — SALVADOR 1986
Podarcis pityusensis gastabiensis — BAUER & GÜNTHER 1995

Podarcis pityusensis gorrae (EISENTRAUT, 1928)
Lacerta lilfordi gorrae EISENTRAUT 1928: 122
Podarcis pityusensis gorrae — SALVADOR 1984
Podarcis pityusensis gorrae — SALVADOR 1986
Podarcis pityusensis gorrae — BAUER & GÜNTHER 1995

Podarcis pityusensis hedwigkamerae (MÜLLER 1927)
Lacerta lilfordi hedwig-kamerae MÜLLER 1927
Podarcis pityusensis hedwigkamerae — SALVADOR 1986: 245
Podarcis pityusensis hedwigkamerae — BERG et al. 2015

Podarcis pityusensis hortae (BUCHHOLZ 1954)
Lacerta pityusensis hortae BUCHHOLZ 1954: 86
Podarcis pityusensis hortae — SALVADOR 1984
Podarcis pityusensis hortae — SALVADOR 1986

Podarcis pityusensis kamerianus (MERTENS 1927)
Lacerta pityusensis kameriana MERTENS 1927: 302
Lacerta lilfordi kameriana — MÜLLER 1927: 194
Lacerta lilfordi zenonis MÜLLER 1928
Lacerta lilfordi zenonis — MERTENS 1934: 83
Lacerta lilfordi kameriana — MERTENS 1934: 71ff
Podarcis pityusensis kameriana — SALVADOR 1984
Podarcis pityusensis kameriana — SALVADOR 1986
Podarcis pityusensis kamerianus — SINDACO & JEREMČENKO 2008

Podarcis pityusensis maluquerorum (MERTENS 1921)
Lacerta lilfordi maluquerorum — MÜLLER 1927: 189
Podarcis pityusensis maluquerorum — SALVADOR 1984
Podarcis pityusensis maluquerorum — SALVADOR 1986
Podarcis pityusensis maluquerorum — SINDACO & JEREMČENKO 2008

Podarcis pityusensis muradae (EISENTRAUT, 1928)
Lacerta lilfordi muradae EISENTRAUT 1928: 122
Lacerta lilfordi muradae — MERTENS 1934: 88
Podarcis pityusensis muradae — SALVADOR 1984
Podarcis pityusensis muradae — SALVADOR 1986
Podarcis pityusensis muradae — BAUER & GÜNTHER 1995

Podarcis pityusensis negrae (EISENTRAUT, 1928)
Lacerta lilfordi negrae EISENTRAUT 1928: 468
Lacerta lilfordi intermedia EISENTRAUT 1928: 123
Lacerta lilfordi negrae EISENTRAUT 1928 (nomen novum)
Lacerta pityusensis negrae — BUCHHOLZ 1954: 81
Podarcis pityusensis negrae — SALVADOR 1984
Podarcis pityusensis negrae — SALVADOR 1986
Podarcis pityusensis negrae — BAUER & GÜNTHER 1995

Podarcis pityusensis ratae (EISENTRAUT, 1928)
Lacerta lilfordi ratae EISENTRAUT 1928: 466
Podarcis pityusensis ratae — SALVADOR 1986
Podarcis pityusensis ratae — BAUER & GÜNTHER 1995

Podarcis pityusensis redonae (EISENTRAUT, 1928)
Lacerta lilfordi redonae EISENTRAUT 1928: 123
Lacerta lilfordi grossae MÜLLER 1929
Podarcis pityusensis redonae — SALVADOR 1984
Podarcis pityusensis redonae — SALVADOR 1986
Podarcis pityusensis redonae — BAUER & GÜNTHER 1995
Podarcis pityusensis redonae — ZAWADZKI 2014

Podarcis pityusensis schreitmuelleri (MÜLLER 1927)
Lacerta lilfordi schreitmülleri MÜLLER 1927: 268
Podarcis pityusensis schreitmuelleri — SALVADOR 1984
Podarcis pityusensis schreitmuelleri — SALVADOR 1986

Podarcis pityusensis tagomagensis (MÜLLER 1927)
Lacerta lilfordi tagomagensis MÜLLER 1927: 267
Lacerta lilfordi tagomagensis — MERTENS 1934: 64
Podarcis pityusensis tagomagensis — SALVADOR 1984
Podarcis pityusensis tagomagensis — SALVADOR 1986

Podarcis pityusensis torretensis (BUCHHOLZ 1954)
Lacerta pityusensis torretensis BUCHHOLZ 1954: 76
Podarcis pityusensis torretensis — SALVADOR 1984
Podarcis pityusensis torretensis — SALVADOR 1986: 247

Podarcis pityusensis vedrae (MÜLLER, 1927)
Lacerta lilfordi vedrae MÜLLER 1927: 190
Lacerta lilfordi vedranellensis MÜLLER 1928
Lacerta lilfordi vedrae — MERTENS 1934: 83
Podarcis pityusensis vedrae — SALVADOR 1984
Podarcis pityusensis vedrae — SALVADOR 1986
Podarcis pityusensis vedrae — KRONIGER & ZAWADZKI 2002
Podarcis pityusensis vedrae — SINDACO & JEREMČENKO 2008
Podarcis pityusensis vedrae — ZAWADZKI 2013
Podarcis pityusensis vedrae — ZAWADZKI 2014 
DistributionSpain (Balearic Islands (Pityuses): Ibiza Island and small, offshore islands (Sal Rossa, Negres, Calders, Punta Sa Ferradura, Bosc, Ses Illetes de Porroig), Penjats Island, S'Illeta de Cala Salada, Can∏ Island, CaragolÚ Island, Conillera Island, Sa Mesquida Island, Formentera, Sabina, Alga, Pouet, Conejo de Formentera, Espalmador, Espardell and Porcs, Island Espardell de Espartar, Gastavâ Island, Islands Na Bosc and Na Gorra, Ses Margalidas Island, Hort Island, Islands Espartar and Escull de S'Espartar, Islands Bleda Plana and Escull Vermell, Murada Island, Negres Island, Rates Island, Islands Santa EulÃria and Redona, Malvâns Islands, Tagomago Island, Torretes Island, Islands Vedrá and Vedranell, introduced to Mallorca)

affinis: Ibiza; Type locality: Malvin Pequeno

ahorcadosi: Type locality: Ahorcados = Penjats

canaretensis: Type locality: Illot d’es Canaret

canensis: Type locality: Canä

caragolensis: Type locality: Caragole

carlkochi: Type locality: “Conichera (Cunillera), westl. Ibiza” [Balearic Islands, Spain]

frailensis: Type locality: Fraile = Espardell de Espartar

gastabiensis: Gastabi

gorrae: Type locality: Bleda Gorra

hortae: Type locality: Hort

kameriana: Esparto Island. Type locality: “Insel Esparto (Espartâ oder Espartal), Westküste von Ibiza, Pityusen.”

maluquerorum: Bledas Island (off Ibiza); Type locality: Bleda

muradae: Type locality: Murada

negrae: Type locality: Negres

pityusensis: Ibiza; Type locality: Ibiza, Pityusen.

ratae: Type locality: Ratas

redonae: Type locality: Redona de Santa Euläria

schreitmülleri: Ibiza; Type locality: Malvin Grande

tagomagensis: Type locality: “Insel Tagomago, östl. von Ibiza” [Balearic Islands, Spain]

torretensis: Type locality: Torretas

vedrae: Es Vedrá Island (near Ibiza, Spain); Type locality: Vedrá Island. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesHolotype: unlocated
Holotype: ZSM (SLM) 1547 (lost), male [affinis]
Holotype:ZSM, Uncatalogued specimen (lost) [calaesaladae]
Holotype: ZSM (SLM) 2226 (lost) [caldesiana]
Holotype: ZFMK 12497, paratypes in ZFMK [caragolensis]
Holotype: ZSM (SLM)1542 (lost), male, collected by C. Koch, 1927 [carlkochi]
Holotype: ZFMK 12467, paratypes in ZFMK and MTKD [characae]
Holotype: ZSM (SLM) 1630 (lost), male [espalmadoris]
Holotype: ZSM (SLM) 1668 (lost), male [grossae]
Holotype: ZSM (SLM) 1632 (lost), male [grueni]
Holotype: ZSM (SLM) 1550 (given as “Nr. 155” in the original description) (lost), male [hedwigkamerae]
Holotype: ZFMK 12410, paratypes in ZFMK and MTKD [hortae]
Holotype: ZFMK 12411, paratypes in ZFMK and MTKD [puercosensis]
Holotype: ZFMK 12409, paratypes in ZFMK and MTKD [purroigensis]
Holotype: ZFMK 12607, paratypes in ZFMK [sabinae]
Holotype: ZSM (SLM) 1544 (lost), male [schreitmuelleri]
Holotype: ZSM (SLM) 1545, male [tagomagensis]
Holotype: ZFMK 12408, paratypes in ZFMK and MTKD [torretensis]
Holotype: ZSM (SLM) 1551 (lost), male [vedrae]
Holotype: ZSM (SLM) 2149 (lost) [vedranellensis]
Holotype: ZSM, uncatalogued (lost) [zenonis] 
CommentSynonymy: Podarcis pityusensis canaretensis CIRER 1980 doesn’t seem to be recognized by most authors but we don’t know of any formal synonymization. CIRER 1987 recognizes only 6 subspecies (as listed by SINDACO & JEREMCENKO 2008). Lacerta lilfordi formenterae MÜLLER 1928 was described independently of Lacerta lilfordi formenterae EISENTRAUT 1928, but both are synonyms. Similarly, Lacerta lilfordi frailensis MÜLLER 1928 is a synonym of Lacerta lilfordi frailensis EISENTRAUT 1928 which was published one month earlier (Franzen & Glaw 2007). BUCHHOLZ 1954 listed gastabiensis as synonym of espalmadoris.

Subspecies: the subspecies indicated as “invalid” are not listed by Salvador 1986.

Types: the types of the Buchholz subspecies are all in the ZFMK (BÖHME 1974). 
Etymologynamed after the type locality. P. p. maluquerorum has been named after Joaquim Maluquer who provided the types to Mertens. P.p. kameriana is named after the collector of the types, Z. Kamer. 
References
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