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Proctoporus sucullucu DOAN & CASTOE, 2003

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common Names 
SynonymProctoporus sucullucu DOAN & CASTOE 2003
Proctoporus sucullucu — CASTOE et al. 2004 
DistributionPeru (Cusco, Apurimac), elevation 3000-3300 m.

Type locality: from Piscacucho (13° 12.213’ S, 72° 22.533’
W), a small village near the town of Chilca, Province of Urubamba, Department of Cusco, Peru; 3191 m elevation.  
Reproductionoviparous 
TypesHolotype: UTA R-51496 
DiagnosisDiagnosis: (1) Nasal divided, forming a pentagonal loreal scale, (2) median occipital present, (3) superciliaries four, first expanded onto dorsal surface of head, (4) palpebral eye-disc made up of a single, undivided, pigmented scale, (5) supralabials six, (6) infralabials five, (7) genials two, (8) dorsal scales quadrangular, with rounded keel, (9) transverse rows of dorsals 38–46, (10) transverse ventral rows 24– 25, (11) a continuous series of small lateral scales separating dorsals and ventrals, (12) femoral pores per hind limb in males 6–8, in females 0–3, (13) preanal pores absent, (14) limbs overlapping when adpressed against the body, (15) ventral scales with grey stippling. Proctoporus sucullucu is a member of the P. pachyurus group (Uzzell, 1970), which is united by the synapomorphy of the presence of an undivided palpebral disc. Proctoporus sucullucu can be distinguished by this charac-ter from all other Proctoporus species except those of the P. pachyurus group. Proctoporus sucullucu can be distinguished from P. boli-vianus by having six supralabials (P. bolivianus has four or five). It can be distinguished from P. guentheri by a higher number of transverse dorsal scale rows (29–36 in P. guentheri; 38–46 in P. sucullucu). It can be distinguished from P. pachyurus by having fewer transverse dorsal scale rows (47–61 in P. pachyurus; 38–46 in P. sucullucu). It can be distinguished from P. unsaacae by limbs overlapping when ad-pressed (from DOAN & CASTOE 2003). 
Comment 
EtymologyNamed after the name of Proctoporus lizards in the local Quechua language. 
References
  • Castoe, T.A.; Doan, T.M. & Parkinson, C.L. 2004. Data partitions and complex models in Bayesian analysis: the phylogeny of Gymnophthalmid lizards. Systematic Biology 53 (3): 448-469 - get paper here
  • Doan, T. M. & Castoe, T.A. 2003. Using morphological and molecular evidence to infer species boundaries within Proctoporus bolivianus Werner (Squamata: Gymnopthalmidae). Herpetologica 59 (3): 432–449 - get paper here
  • Doan, T. M., Sheffer, S. A., Warmington, N. R., & Evans, E. E. 2021. Population biology of the unusual thermoconforming lizards of the Andes Mountains of Peru (Squamata: Gymnophthalmidae). Austral Ecology - get paper here
  • Doan, T.F.; Castoe, T.A. & Arizábal Arriaga, W. 2005. Phylogenetic relationships of the genus Proctoporus sensu stricto (Squamata: Gymnophthalmidae), with a new species from Puno, Southeastern Peru. Herpetologica 61 (3): 325-336 - get paper here
  • Goicoechea, Noemí; José M. Padial, Juan C. Chaparro, Santiago Castroviejo-Fisher, Ignacio De la Riva 2012. Molecular phylogenetics, species diversity, and biogeography of the Andean lizards of the genus Proctoporus (Squamata: Gymnophthalmidae). Molecular Phylogenetics and Evolution, 65 (3): 953–964 [print version published in 2013], http://dx.doi.org/10.1016/j.ympev.2012.08.017
  • Köhler, G. and E. Lehr 2004. Comments on Euspondylus and Proctoporus (Squamata: Gymnophthalmidae) from Peru, with the description of three new species and a key to the Peruvian species. Herpetologica 60 (4):501-518 - get paper here
 
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