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Proctoporus unsaacae DOAN & CASTOE, 2003

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards) 
Subspecies 
Common Names 
SynonymProctoporus unsaacae DOAN & CASTOE 2003
Proctoporus unsaacae — CASTOE et al. 2004 
DistributionPeru (Cusco)

Type locality: from Quello Uno (13° 21.887’ S, 71° 58.215’ W), a village near the town of Calca, Province of Calca, Department of Cusco, Peru; 3253 m elevation.  
Reproductionoviparous 
TypesHolotype: UTA R-51488 
DiagnosisDiagnosis. (1) Nasal divided, forming a pentagonal loreal scale; (2) median occipital present; (3) superciliaries four, first expanded onto dorsal surface of head; (4) palpebral eye-disc made up of a single, undivided scale; (5) genials two; (6) dorsal scales quadrangular, with low rounded keel, surrounded or not by longitudinal striations; (7) transverse rows of dorsals 36–42; (8) transverse ventral rows 18– 26; (9) a continuous series of small lateral scales separating dorsals and ventrals; (10) femoral pores per hind limb in males 5–7, in females 0–3; (11) preanal pores absent; (12) limbs not overlapping when adpressed against the body in adults; (13) a continuous series of dark-centered ocelli on the lateral surface; (14) venter cream, medial scales immaculate, ventrolateral scales with dark spot centered in each scale. Specimens of P. unsaacae show the pres-ence of an undivided palpebral eye-disc, which place them in the P. pachyurus group (Uzzell, 1970). Proctoporus unsaacae can be distin-guished from all other Proctoporus species except those in the P. pachyurus group by this character. Proctoporus unsaacae can be distinguished from P. bolivianus by a continuous series of lateral ocelli (in P. bolivianus, ocelli, if present, are faint and do not form a continuous series). It can be distinguished from P. guentheri by cream venter with ventrolateral scales that each have a dark spot (P. guentheri has yellow or orange venter without any spots). It can be distinguished from P. pachyurus by a lower number of transverse dorsal scale rows (47–61 in P. pachyurus; 36–42 in P. unsaacae). It can be distinguished from P. sucullucu by limbs not overlapping when adpressed. 
Comment 
EtymologyNamed in honor of the herpetological research group at the Universidad Nacional de San Antonio Abad de Cusco, a university in Cusco, Peru, commonly referred to as UNSAAC. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Castoe, T.A.; Doan, T.M. & Parkinson, C.L. 2004. Data partitions and complex models in Bayesian analysis: the phylogeny of Gymnophthalmid lizards. Systematic Biology 53 (3): 448-469 - get paper here
  • Doan, T. M. & Castoe, T.A. 2003. Using morphological and molecular evidence to infer species boundaries within Proctoporus bolivianus Werner (Squamata: Gymnopthalmidae). Herpetologica 59 (3): 432–449 - get paper here
  • Doan, T.F.; Castoe, T.A. & Arizábal Arriaga, W. 2005. Phylogenetic relationships of the genus Proctoporus sensu stricto (Squamata: Gymnophthalmidae), with a new species from Puno, Southeastern Peru. Herpetologica 61 (3): 325-336 - get paper here
  • Goicoechea, Noemí; José M. Padial, Juan C. Chaparro, Santiago Castroviejo-Fisher, Ignacio De la Riva 2013. Molecular phylogenetics, species diversity, and biogeography of the Andean lizards of the genus Proctoporus (Squamata: Gymnophthalmidae). Molecular Phylogenetics and Evolution, Volume 65, Issue 3, December 2012, Pages 953–964 - get paper here
  • Köhler, G. and E. Lehr 2004. Comments on Euspondylus and Proctoporus (Squamata: Gymnophthalmidae) from Peru, with the description of three new species and a key to the Peruvian species. Herpetologica 60 (4):501-518 - get paper here
 
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