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Protobothrops kaulbacki (SMITH, 1940)

IUCN Red List - Protobothrops kaulbacki - Data Deficient, DD

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Higher TaxaViperidae, Crotalinae, Colubroidea, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesE: Kaulback’s lance-headed pitviper
G: Kaulbacks Grubenotter 
SynonymTrimeresurus kaulbacki SMITH 1940: 485
Trimeresurus kaulbacki — SMITH 1943: 512
Trimeresurus kaulbacki — WELCH 1994: 117
Protobothrops kaulbacki — KRAUS et al. 1996
Protobothrops kaulbacki — LEVITON et al. 2003
Protobothrops kaulbacki — GUMPRECHT et al. 2004
Protobothrops kaulbacki — WALLACH et al. 2014: 572 
DistributionN Myanmar (= Burma), China, India (Arunachal Pradesh, Papum Pare District)

Type locality: Pangnamdim, north of the Triangle, Upper Burma. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous. 
TypesHolotype: BMNH 1946.1.19.23–24); 1943:512 
CommentVenomous!

DIAGNOSIS (DIAGNOSTIC CHARACTERS).— Scales in 23–25 longitudinal rows at midbody; body elongate, head long, massive, with narrow snout; canthus rostralis sharp; single large, squarish loreal; first upper labial completely separated from nasal by a suture; second upper labial anteriorly bordering facial pit; 8–10 scales in line between supraoculars; supraoculars usually single, flat, without a vertical projection; dull grayish or olive green with large, blackish rhombohedral dorsal blotches, either distinct or united to one another; smaller spots on sides; symmetrical yellow lines on head; subcaudals paired; ventrals 201–212; subcaudals 66–78, some of the anterior scutes may be single, others paired; hemipenes with spines. Total length males 1340 mm, females 1410 mm; tail length males 225 mm, females 230 mm. [after LEVITON 2003]

Distribution: see map in Guo et a. 2016: 383 (Fig. 1). 
References
  • Bhide, K.S.; Captain, A.; Bhatt, B.B.; Gumprecht, A. & Tillack, F. 2008. Protobothrops kaulbacki (SMITH). Sauria 30 (2): 2 - get paper here
  • Dowling, H.G., & Jenner, J.V. 1988. Snakes of Burma: checklist of reported species and bibliography. Smithsonian Herp. Inf. Serv. (76): 19 pp. - get paper here
  • Gumprecht, A.; Tillack, F.; Orlov, N.L.; Captain, A. & Ryabow, S. 2004. Asian pitvipers. Geitje Books, Berlin, 368 pp.
  • Guo, Peng, Anita Malhotra, Pi P. Li, Catharine E. Pook and Simon Creer. 2007. New evidence on the phylogenetic position of the poorly known Asian pitviper Protobothrops kaulbacki (Serpentes: Viperidae: Crotalinae) with a redescription of the species and a revision of the genus Protobothrops. The Herpetological Journal 17 (4):237-247
  • GUO, PENG; QIN LIU, TAO WEN, RONG XIAO, MING FANG, GUANGHUI ZHONG, NGUYEN Q. TRUONG, FEI ZHU, ROBERT C. JADIN, CAO LI 2016. Multilocus phylogeny of the Asian Lance-headed pitvipers (Squamata, Viperidae, Protobothrops) Zootaxa 4093 (3): 382–390 - get paper here
  • Leviton, Alan E.; Guinevere O.U. Wogan; Michelle S. Koo; George R. Zug; Rhonda S. Lucas and Jens V. Vindum 2003. The Dangerously Venomous Snakes of Myanmar Illustrated Checklist with Keys. Proc. Cal. Acad. Sci. 54 (24): 407–462
  • Orlov, Nikolai L.; Sergei A. Ryabov, and Nguyen Thien Tao 2009. Two New Species of Genera Protobothrops Hoge et Romano-Hoge, 1983 and Viridovipera Malhotra et Thorpe, 2004 (Ophidia: Viperidae: Crotalinae) from Karst Region in Northeastern Vietnam. Part I. Description of a New Species of Protobothrops Genus. Russ. J. Herpetol. 16 (1): 69-82 - get paper here
  • PAN, Hujun; Basundhara CHETTRI, Daode YANG, Ke JIANG, Kai WANG, Liang ZHANG and Gernot VOGEL 2013. A New Species of the Genus Protobothrops (Squamata: Viperidae) from Southern Tibet, China and Sikkim, India. Asian Herpetological Research 4 (2): 109–115 - get paper here
  • Rao D. & Zhao E. 2005. A NEW RECORD FROM CHINA-PROTOBOTHROPS KAULBACKI (REPTILIA, SERPENTES, VIPERIDAE). ACTA ZOOTAXONOMICA SINICA 2005 30 (1), 209 - 211
  • Smith, M.A. 1940. The Amphibians and Reptiles obtained by Mr. Ronald Kaulback in Upper Burma. Records of the Indian Museum 42: 465-486
  • Smith, M.A. 1943. The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp.
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, 1237 pp.
  • YANG, JIAN-HUAN; NIKOLAI L. ORLOV & YING-YONG WANG 2011. A new species of pitviper of the genus Protobothrops from China (Squamata: Viperidae). Zootaxa 2936: 59–68 - get paper here
  • Zhao, E.M. 2006. The snakes of China [in Chinese]. Hefei, China, Anhui Sience & Technology Publ. House, Vol. I, 372 pp., Vol. II (color plates), 280 pp.
 
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