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Pseudogekko sumiklab SILER, DAVIS, WATTERS, FREITAS, GRIFFITH, BINADAY, LOBOS AMARGA & BROWN, 2017

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesE: Bicol Gecko 
SynonymPseudogekko sumiklab SILER, DAVIS, WATTERS, FREITAS, GRIFFITH, BINADAY, LOBOS AMARGA & BROWN 2017 
DistributionPhilippine Islands (Luzon)

Type locality: foothills of Mt. Mayon, Sitio Nagsipit, Barangay Mariroc, Municipality of Tabaco, Albay Province, Luzon Island, Philippines (13.305588N, 123.688988E; in all cases, datum 1⁄4 WGS84; 399 m elev.),  
Reproduction 
TypesHolotype: PNM 9843 (DRD Field No. 2700, formerly OMNH 44874), adult female, collected between 1900 and 2200 h on 5 March 2016, by CDS, DRD, OWG, N. Huron, J. Fernandez, J. Bulalacao, W. Bulalacao, and B. Gurobat. Paratype. KU 343847 (RMB 22872), adult female, collected 2040 h on 27 January 2017, on Mt. Cawayan, Municipality of Irosin, Sorsogon Province, Luzon Island, Philippines (12.701418N, 124.078208E; 280 m elev.), by JWBB, J. Fernandez, and RMB. 
DiagnosisDiagnosis. Pseudogekko sumiklab can be distinguished from congeners by the following combination of characters: (1) body size small (SVL 41.8, 46.6 mm [holotype, paratype]); (2) axilla–groin distance short (23.6, 25.8 mm); (3) head length short (5.8, 7.1 mm); (4) absolute snout length short (4.0, 5.1 mm); (5) relative snout length long (64.7, 69.2% HL); (6) Finger-III scansors 11, 12; (7) Toe-IV scansors 13, 14; (8) supralabials 14; (9) infralabials 13, 14; (10) circumorbitals 39, 46; (11) paravertebrals 220, 224; (12) ventrals 106, 109; (13) enlarged precloacals 13; (14) femoral pore-bearing scales absent; (15) cephalic spots present, black and cream; (16) dorsolateral trunk spots present; (17) black vertebral spots present; (18) limb spots absent; (19) transverse tail bands absent; (20) trunk stripes absent; (21) interorbital band absent; (22) iris ring coloration absent; and (23) significant genetic divergence (Tables 2–4; Figs. 2–4).

Comparisons.—Several characters distinguish Pseudo- gekko sumiklab from all other species of Pseudogekko (Tables 3, 4). Pseudogekko sumiklab most closely resembles P. atiorum and P. brevipes; however, it differs from these species in several important characters. Pseudogekko sumiklab can be distinguished from P. atiorum by having a shorter absolute head length (5.8, 7.1 mm vs. 7.4–9.4 mm) and head width (5.6, 5.7 mm vs. 5.9–8.3 mm) and fewer supralabials (14 vs. 15–17), paravertebrals (220, 224 vs. 226– 240), and ventrals (106, 109 vs. 119–129). The new species further differs from this taxon by a greater number of circumorbitals (39, 46 vs. 35–38), the presence (vs. absence) of vertebral spots, the absence (vs. presence) of an interorbital band, and the presence (vs. absence) of a subcaudal stripe. From P. brevipes the new species differs by having a greater total length (87.8, 91.0 mm vs. 72.0–87.5 mm), fewer infralabials (13, 14 vs. 14, 15), more circum- orbitals (39, 46 vs. 33–35), paravertebrals (220, 224 vs. 211– 218), and enlarged precloacal pore-bearing scales (13 vs. 12) and the absence (vs. presence) of an interorbital band and presence (vs. absence) of a subcaudal stripe.
Pseudogekko sumiklab can be distinguished from P. chavacano by having a smaller body size (SVL ? 46.6 mm vs. 54.7 mm), axilla–groin distance (?25.8 mm vs. 26.7–30.0 mm), head length (?7.8 mm vs. 10.4 mm), head width (?6.7 mm vs. 7.8–8.5 mm), and snout length (4.0 mm vs. 5.8–6.0 mm), and fewer Finger-III scansors (11, 12 vs. 15, 16), Toe- IV scansors (13, 14 vs. 17, 20), supralabials (14 vs. 15, 16), infralabials (13, 14 vs. 16, 17), ventrals (106, 109 vs. 122, 123), and numbers of enlarged, precloacal pore-bearing scales (13 vs. 16). Additionally, the new species has more paravertebrals (220, 224 vs. 195, 197) and is further distinguished by the presence of black and cream (vs. dense neon green) cephalic spots, presence of cream (vs. neon green) dorsolateral spots, presence (vs. absence) of black vertebral spots, absence (vs. presence) of dense neon green limb spots, and absence (vs. presence) of transverse tail bands. From P. compresicorpus, the new species differs by having a smaller body size (SVL ? 46.6 mm vs. 54.9–59.7 mm), axilla–groin distance (?25.8 mm vs. 27.0–32.6 mm), total length (?91.0 mm vs. 105.9–117.3 mm), head length (?7.8 mm vs. 9.2–11.3 mm), head width (?6.7 mm vs. 7.5– 10.1 mm), and snout length (4.0 mm vs. 5.3–7.4 mm), and fewer Finger-III scansors (11, 12 vs. 15–17), Toe-IV scansors (13, 14 vs. 18, 19), supralabials (14 vs. 16–20), paravertebrals (220, 224 vs. 226–234), and ventrals (106, 109 vs. 127–130), and by the presence of sparse black and cream (vs. dense neon green) cephalic spots, presence of cream (vs. faint neon green) dorsolateral spots, presence (vs. absence) of black vertebral spots, absence (vs. presence) of faint neon green limb spots, and absence (vs. presence) of light blue iris ring coloration; from P. ditoy and P. isapa by having a smaller body size (SVL ? 46.6 mm vs. 52.6 mm [P. ditoy], 62.1 mm [P. isapa]), axilla–groin distance (?25.8 mm vs. 25.1–29.7 mm [P. ditoy], 32.1–33.0 mm [P. isapa]), head length (?7.8 mm vs. 9.3–9.6 mm [P. ditoy], 11.7–11.8 mm [P. isapa]), head width (?6.7 mm vs. 7.7–7.9 mm [P. ditoy], 7.4–10.1 mm [P. isapa]), and snout length (4.0 mm vs. 5.4–5.7 mm [P. ditoy], 7.2–7.5 mm [P. isapa]), and fewer Finger-III scansors (11, 12 vs. 14, 15 [P. ditoy], 13 [P. isapa]), Toe-IV scansors (13 vs. 16, 17 [P. ditoy], 17 [P. isapa]), supralabials (14 vs. 17, 20 [P. ditoy], 20, 21 [P. isapa]), infralabials (13, 14 vs. 16, 17 [P. ditoy], 17–19 [P. isapa]), ventrals (106, 109 vs. 111–118 [P. ditoy], 135–141 [P. isapa]), and enlarged precloacal pores (13 vs. 18 [P. ditoy], 15 [P. isapa]), and by the presence (vs. absence) of dorsolateral and vertebral spots; additionally from P. ditoy by having more paravertebrals (220, 224 vs. 180–185) and by the presence (vs. absence) of cephalic spots; additionally from P. isapa the new species differs by having fewer circumorbitals (?46 vs. 50–54) and paravertebrals (220, 224 vs. 240–246), and by the absence (vs. presence) of an interorbital band; from P. pungkaypinit and P. smaragdinus by having a smaller body size (SVL ? 46.6 mm vs. 75.2–75.3 mm [P. pungkaypinit], 50.2–61.7 mm [P. smar- agdinus]), axilla–groin distance (?25.8 mm vs. 37.2–41.2 mm [P. pungkaypinit], 26.2–35.5 mm [P. smaragdinus]), total length (?91.0 mm vs. 125.3–141.2 mm [P. pungkaypinit], 103.6–129.7 mm [P. smaragdinus]), head length (5.8 mm vs. 11.4–13.6 mm [P. pungkaypinit], 7.8–10.7 mm [P. smar- agdinus]), head width (? 6.7 mm vs. 9.3–11.2 mm [P. pungkaypinit], 7.1–9.8 mm [P. smaragdinus]), and snout length (4.0 mm vs. 6.7–7.5 mm [P. pungkaypinit], 4.9–6.5 mm [P. smaragdinus]), and fewer Finger-III scansors (11, 12 vs. 15–17 [P. pungkaypinit], 15–18 [P. smaragdinus]), Toe- IV scansors (13, 14 vs. 17–21 [P. pungkaypinit], 16–22 [P. smaragdinus]), supralabials (14 vs. 16–20 [P. pungkaypinit], 16–19 [P. smaragdinus]), infralabials (13, 14 vs. 17–19 [P. pungkaypinit], 14–17 [P. smaragdinus]), paravertebrals (220, 224 vs. 265–280 [P. pungkaypinit], 241–252 [P. smaragdi- nus], ventrals (106, 109 vs. 125–155 [P. pungkaypinit], 124– 130 [P. smaragdinus]), enlarged pore-bearing scales (13 vs. 17–20 [P. pungkaypinit], 32–41 [P. smaragdinus], and by the presence (vs. absence) of vertebral spots; additionally from P. pungkaypinit by having fewer circumorbitals (? 46 vs. 50–55), the presence (vs. absence) of cephalic spots, presence (vs. absence) of dorsolateral spots, and absence (vs. presence) of trunk stripes; additionally from P. smaragdinus by having a greater number of circumorbitals (?39 vs. 33– 35), the absence (vs. presence) of femoral pores, absence (vs. presence) of bright neon yellow to orange (undisturbed) to neon green (disturbed) body coloration, and absence (vs. presence) of transverse tail bands. 
Comment 
EtymologyThe authors derived the masculine specific epithet from the Tagalog (Filipino) verb sumiklab, meaning to burst out quickly, or hotly, or to ignite and flare up, in reference to the new species type locality, Mt. Mayon, and the other volcanos of the Bicol Peninsula. One of the world’s most active and dangerous volcanoes, Mayon’s near-perfect, symmetrical cone is an iconic feature of the Bicol Region volcanic landscape and a picturesque symbol of the national significance of Bicolandia in Philippine culture (Fig. 5). 
References
  • Siler, Cameron D.; Drew R. Davis, Jessa L. Watters, Elyse S. Freitas, Oliver W. Griffith, Jake Wilson B. Binaday, Athena Heart T. Lobos, Ace Kevin S. Amarga, and Rafe M. Brown 2017. First Record of the Pseudogekko brevipes Complex from the Northern Philippines, with Description of a New Species Herpetologica 73 (2): 162-175 - get paper here
 
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