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Pseudotrapelus chlodnickii MELNIKOV, ŚMIEŁOWSKI, MELNIKOVA, NAZAROV & ANANJEVA, 2015

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common Names 
SynonymPseudotrapelus chlodnickii MELNIKOV, ŚMIEŁOWSKI, MELNIKOVA, NAZAROV & ANANJEVA 2015 
DistributionSudan (Jumhūriyyat), Libya

Type locality: Gamamiya, Bayuda Desert, Sudan,
Africa (19°27’18.3’’ N 32°48’40.5’’ E).  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ZISP 28540, subadult male collected on 25 of November 2007 by Jan ŚMIEŁOWSKI (Fig. 2). 
DiagnosisDiagnosis: A large, short legged, chocolate-red colored species of Pseudotrapelus with white ocelli on the dorsum (up to 8 dorsal scales per white ocelli, each white ocelli with dark chocolate red marbling), with the 3rd toe longer than the 4th, with 16 lamellae under the 4th toe, with 6 unseparated well developed precloacal pores in males, and with no enlarged scales in the occipital area of the head and dorsum. 
CommentDistribution: see maps in TAMAR et al. 2016: Fig. 1, Tamar et al. 2019: Fig. 1, Tamar et al. 2023: 1034 (Fig. 1). 
EtymologyThe new species is named in honor of distinguished Polish archeologist Dr. Marek Chłodnicki, the Chief of the Polish Archeological Expedition of the International Archeological Mission in Sudan during archeological excavations before opening the new dam in IV cataract of the Nile River (northern Sudan). The only archeologist who respect present biological field study survey of the recent fauna of this unique region and accept to participate in archeological team also zoologist Jan Śmiełowski. 
References
  • Bauer, Aaron M.; Jonathan C. DeBoer , Dylan J. Taylor 2017. Atlas of the Reptiles of Libya. Proc. Cal. Acad. Sci. 64 (8): 155-318 - get paper here
  • Melnikov, Daniel; Jan Śmiełowski, Ekaterina Melnikova, Roman Nazarov, Natalia B. Ananjeva 2015. Red’n’blues: A New Species of Pseudotrapelus (Agamidae, Sauria) from Sudan, Africa. Russ. J. Herpetol. 22 (1): 53-60 - get paper here
  • Melnikova, Ekaterina; Daniel Melnikov, Natalia Ananjeva 2015. Mitochondrial and Nuclear Gene Phylogeny of Pseudotrapelus (Sauria: Agamidae) with Considerations on the Hybridization and mtDNA Introgression between Species. Russ. J. Herpetol. 22 (1): 61-67 - get paper here
  • Milto, Konstantin D.; Samy A. Saber, Abdullah M. Nagy, Roman A. Nazarov, Daniel A. Melnikov, Natalia B. Ananjeva 2019. First Report on the Reptile Diversity of Wadi El Gemal National Park, Eastern Desert, Egypt. Russ. J. Herpetol. 26 (3): 175-184 - get paper here
  • Spawls, Stephen; Tomáš Mazuch& Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Tamar, K., Chirio, L., Shobrak, M., Busais, S. & Carranza, S. 2019. Using multilocus approach to uncover cryptic diversity within Pseudotrapelus lizards from Saudi Arabia. Saudi J. Biol. Sci. (2019). doi: - get paper here
  • Tamar, K., Uvizl, M., Shobrak, M., Almutairi, M., Busais, S., AlGethami, R. H. M., ... & Šmíd, J. 2023. A new species of Pseudotrapelus (Reptilia: Squamata: Agamidae) from Central Arabia. Vertebrate Zoology 73, 1033-1045 - get paper here
  • Tamar, Karin; Sebastian Scholz, Pierre-André Crochet, Philippe Geniez, Shai Meiri, Andreas Schmitz, Thomas Wilms, Salvador Carranza 2016. Evolution around the Red Sea: Systematics and biogeography of the agamid genus Pseudotrapelus (Squamata: Agamidae) from North Africa and Arabia. Molecular Phylogenetics and Evolution 97 (2016) 55–68 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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