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Ptyas carinata (GÜNTHER, 1858)

IUCN Red List - Ptyas carinata - Least Concern, LC

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesE: Keeled Rat Snake
G: Gekielte Rattenschlange
Chinese: 黑网乌梢蛇 
SynonymCoryphodon carinatus GÜNTHER 1858: 112
Zaocys carinatus GÜNTHER 1864
Zaocys (Zapyrus) carinatus — BOETTGER 1887
Zaocys tenasserimensis SCLATER 1891: 238
Zaocys carinatus — BOULENGER 1893: 377
Ptyas carinatus — TAYLOR 1922: 136
Ptyas carinatus — WALL 1923
Zaocys carinatus — SMITH 1943
Zaocys carinatus — GRANDISON 1978: 291
Ptyas carinatus — DAVID & VOGEL 1996
Zaocys carinatus — MANTHEY & GROSSMANN 1997: 402
Ptyas carinatus — COX et al. 1998: 55
Zaocys carinatus — GRISMER et al. 2002
Zaocys carinatus — ROOIJEN & ROOIJEN 2002
Ptyas carinatus — MALKMUS et al. 2002
Ptyas carinata — DAVID & DAS 2004
Zaocys carinatus — ZHAO 2006
Ptyas carinata — WALLACH et al. 2014: 615 
DistributionBurma (Myanmar), China (Yunnan),
Indonesia (Sumatra, Java, Borneo), Laos,
Malaysia (Malaya and East Malaysia, Pulau Tioman),
Philippines (Palawan), Singapore Island,
Thailand, Vietnam, Cambodia

Type locality: Borneo  
Reproductionoviparous. 
TypesHolotype: BMNH (presumed, status unclear fide P. Campbell, pers. comm., March 2019). 
Diagnosis 
CommentHabitat: partly arboreal (Harrington et al. 2018). 
Etymology 
References
  • Bergman, R.A.M. 1952. L'anatomie du genre Ptyas a Java. Riv. Biol. Colon., Roma 12:1-42.
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • Das, I. 2012. A Naturalist's Guide to the Snakes of South-East Asia: Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. Oxford J, ohn Beaufoy Publishing - get paper here
  • David, P. & I. Das 2004. On the grammar of the gender of Ptyas Fitzinger, 1843 (Serpentes: Colubridae). Hamadryad 28 (1&2): 113-116
  • David,P. & Vogel, G. 1996. The snakes of Sumatra. An annotated checklist and key with natural history notes. Bücher Kreth, Frankfurt/M.
  • Dowling, H.G., & Jenner, J.V. 1988. Snakes of Burma: checklist of reported species and bibliography. Smithsonian Herp. Inf. Serv. (76): 19 pp. - get paper here
  • Grandison, A. G. C. 1978. Snakes of West Malaysia and Singapore. Annalen des Naturhistorischen Museums in Wien 81 [1977]: 283-303 - get paper here
  • Grismer, L.L. 2011. Amphibians and reptiles of the Seribuat Archipelago. Edition Chimaira, Frankfurt, 239 pp.
  • Grismer, L.L., McGuire, J.A.; Sosa, R. & Kaiser, H. 2002. Revised checklist and comments on the terrestrial herpetofauna of Pulau Tioman, Peninsular Malaysia. Herpetological Review 33 (1): 26-29 - get paper here
  • Günther, A. 1858. Catalogue of Colubrine snakes of the British Museum. London, I - XVI, 1 - 281
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Lidth DE JEUDE, T.W. VAN 1922. Snakes from Sumatra. Zoologische Mededelingen 6: 239-253. - get paper here
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Rooijen, J. van & van Rooijen, M. 2002. Einige Ergänzungen, Berichtigungen und neue Beobachtungen zur Herpetofauna von Pulau Tioman, West-Malaysia. Sauria 24 (3): 3-12 [erratum in 24 (4): 34] - get paper here
  • SACHA, M. 2015. Herpetoreisen in Thailand: Erste Eindrücke aus Krabi. Sauria 37 (3): 43-55 - get paper here
  • Sclater,W.L. 1891. Notes on a collection of snakes in the Indian Museum, with descriptions of several new species. J. Asiat. Soc. Bengal LX: 230-250 - get paper here
  • Smith, M.A. 1943. The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp.
  • Stuart, B.; Sok, K. & Neang, T. 2006. A collection of amphibians and reptiles from hilly Eastern Cambodia. Raffles Bull. Zool. 54 (1): 129-155 - get paper here
  • Taylor, E.H. 1922. The snakes of the Philippine Islands. Manila (Bureau of Printing or Science), Monograph 16: 312 pp. - get paper here
  • Taylor,E.H. 1965. The serpents of Thailand and adjacent waters. Univ. Kansas Sci. Bull. 45 (9): 609-1096 - get paper here
  • Teo, R.C.H. & N.J. Thomas 2019. Updated Inventory of Amphibians, Reptiles and Mammals of the Bukit Timah Nature Reserve, Singapore. Gardens’ Bulletin Singapore 71(Suppl. 1):145-183; doi: 10.26492/gbs71(suppl.1).2019-06 - get paper here
  • Teo, R.C.H. & Rajathurai, S. 1997. Mammals, reptiles and amphibians in the Nature Reserves of Singapore - diversity, abundance and distribution. Proc. Nature Reserves Survey Seminar. Gardens’ Bulletin Singapore 49: 353-425
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Weinell, Jeffrey L.; Errol Hooper, Alan E. Leviton, Rafe M. Brown 2019. Illustrated Key to the Snakes of the Philippines. Proc. Cal. Acad. Sci. (4) 66 (1): 1-49 - get paper here
  • Zhao, E.M. 2006. The snakes of China [in Chinese]. Hefei, China, Anhui Sience & Technology Publ. House, Vol. I, 372 pp., Vol. II (color plates), 280 pp.
 
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