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Ptyas major (GÜNTHER, 1858)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Common NamesE: Chinese Green Snake
G: Chinesische Grünnatter, Chinesische Grasnatter
V: Ran dai lon
Chinese: 翠青蛇 
SynonymCyclophis major GÜNTHER 1858: 120
Cyclophis major — GÜNTHER 1864: 230
Herpetodryas chloris HALLOWELL 1861: 503
Cyclophis major — GÜNTHER 1888: 170
Ablabes major — BOETTGER 1894: 140
Ablabes major — BOULENGER 1894: 279
Entechinus major — COPE 1895
Liopeltis major — STEJNEGER 1907: 338
Coluber delacouri SMITH 1930: 681
Liopeltis major bicarinata MAKI 1931
Eurypholis major — POPE 1935
Opheodrys major — SMITH 1943: 178
Entechinus major — CUNDALL 1981
Liopeltis major — WELCH 1988
Entechinus major — WELCH 1988
Cyclophiops major — OTA 1991
Cyclophiops major — ZIEGLER et al. 2006
Cyclophiops major — WALLACH et al. 2014: 202
Ptyas major — FIGUEROA et al. 2016 
DistributionS China (Hong, Kong, Hainan, northward to Henan and Gansu),
Taiwan, N Vietnam, Laos

Type locality: Ningpo, China  
TypesType: unknown (Sang et al. 2009)
Holotype: NSM H02569 [bicarinata]
Holotype: USNM 7335 [Herpetodryas chloris] 
CommentType species: Entechinus major is the type species of the genus Entechinus COPE 1895.

Behavior: diurnal.

Synonymy: partly fide WALLACH et al. 2014: 200.

Characteristic features. One loreal, eight supralabials (4-5 in contact with the eye), 15 scales across the midbody, 169 ventral scales, anal divided, and 76 divided subcaudal scales (from ZIEGLER et al. 2006).

Habitat: partly arboreal (Harrington et al. 2018). 
  • Barbour, Thomas 1909. Notes on Amphibia and Reptilia from Eastern Asia. Proc. New England zool. Club 4: 53-78, 2 plates - get paper here
  • Boettger, O. 1894. Materialien zur herpetologischen Fauna von China. III. Liste der von der Insel Hainan bekannten Kriechtiere. Ber. Senckenberg. Naturf. Ges.: 129-152 - get paper here
  • Boulenger, George A. 1894. Catalogue of the snakes in the British Museum (Natural History). Volume II., Containing the Conclusion of the Colubridæ Aglyphæ. British Mus. (Nat. Hist.), London, xi, 382 pp. - get paper here
  • Deuve, J. 1961. Liste annotee des Serpents du Laos. Bull. Soc. Sci. Nat. Laos 1:5-32.
  • Figueroa A, McKelvy AD, Grismer LL, Bell CD, Lailvaux SP 2016. A Species-Level Phylogeny of Extant Snakes with Description of a New Colubrid Subfamily and Genus. PLoS One 11 (9): e0161070.doi:10.1371/journal.pone.0161070 - get paper here
  • Günther, A. 1858. Catalogue of Colubrine snakes of the British Museum. London, I - XVI, 1 - 281
  • Günther, A. 1864. The Reptiles of British India. London (Taylor & Francis), xxvii + 452 pp. - get paper here
  • Günther, A. 1888. On a collection of reptiles from China. Ann. Mag. nat. Hist. (6) 1: 165-172 - get paper here
  • Hallowell, E. 1861. Report upon the Reptilia of the North Pacific Exploring Expedition, under command of Capt. John Rogers, U. S. N. Proc. Acad. Nat. Sci. Philadelphia 12 [1860]: 480 - 510 - get paper here
  • Kunz, Kriton 2015. Schlangen, die Wirbellose fressen. Terraria-Elaphe 2015 (5): 14-20 - get paper here
  • Maki, M. 1931. Monograph of the Snakes of Japan. Dai-ichi Shobo, Tokyo. (1), 7, 240 pages (expanded edition 1933)
  • Mell,R. 1931. List of Chinese snakes. Lingnan Sci. Jour., Canton, 8 [1929]: 199-219.
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Smith, M.A. 1930. Two new Snakes from Tonkin, Indo-China. Ann. Mag. nat. Hist. (10) 6: 681-683
  • Smith, M.A. 1943. The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp.
  • Stejneger, L. 1910. The batrachians and reptiles of Formosa. Proc. US Natl. Mus. 38: 91-114 - get paper here
  • Stejneger, LEONHARD H. 1907. Herpetology of Japan and adjacent territory. Bull. US Natl. Mus. 58: xx, 1-577 - get paper here
  • Toriba, M. 1993. Present location of the type specimens described by Maki. The Snake 25: 71-72
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Zhao, E.M. 2006. The snakes of China [in Chinese]. Hefei, China, Anhui Sience & Technology Publ. House, Vol. I, 372 pp., Vol. II (color plates), 280 pp.
  • Zhao,E. & Adler,K. 1993. Herpetology of China. SSAR, Oxford/Ohio, 1-522
  • Ziegler, Thomas; Annemarie Ohler; Vu Ngoc Thanh; Le Khac Quyet; Nguyen Xuan Thuan; Dinh Huy Tri & Bui Ngoc Thanh 2006. Review of the amphibian and reptile diversity of Phong Nha – Ke Bang National Park and adjacent areas, central Truong Son, Vietnam. Herpetologica Bonnensis II: 247-262 - get paper here
  • ZIEGLER, THOMAS; RALF HENDRIX, VU NGOC THANH, MARTINA VOGT, BERNHARD FORSTER & DANG NGOC KIEN 2007. The diversity of a snake community in a karst forest ecosystem in the central Truong Son, Vietnam, with an identification key. Zootaxa 1493: 1-40 - get paper here
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