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Rena segrega (KLAUBER, 1939)

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Higher TaxaLeptotyphlopidae, Epictinae, Epictini, Typhlopoidea, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Trans-Pecos Blind Snake/Threadsnake
G: Trans-Pecos Schlankblindschlange
S: Culebra Lombriz de Trans- Pecos 
SynonymLeptotyphlops humilis segregus KLAUBER 1939
Leptotyphlops humilis segregus — JAMESON & FLURY 1949
Leptotyphlops humilis segregus — MILSTEAD 1953
Leptotyphlops humilis chihuahuaensis TANNER 1985: 623
Leptotyphlops humilis segregus — CONANT & COLLINS 1991: 145
Leptotyphlops humilis segregus — CROTHER 2000: 65
Leptotyphlops humilis segregus — TENNANT & BARTLETT 2000: 34
Rena humilis segregus — CROTHER et al. 2012
Rena segrega — WALLACH et al. 2014: 631
Rena humilis segrega — CROTHER et al. 2017
Rena segrega — VILLELA-FLORES et al. 2022 
DistributionUSA (SE Arizona, Texas [Trans-Pecos], New Mexico), Mexico (San Luis Potosí, N Coahuila, Nuevo León, Durango, Chihuahua)

Type locality: Chalk Draw, Brewster County, Texas  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: USNM 103670 
Diagnosis 
CommentSynonymy: chihuahuaensis has been considered as a synonym of humilis, but synonymized with segrega by Lemos-Espinal and Smith (2007) and Flores-Villela et al. 2022. 
EtymologyThe specific name is a Latin word segrego meaning "separate," and was applied to the population in western Texas, which was isolated from other populations of Rena humilis, as then understood (LEMOS-ESPINAL & DIXON 2013). 
References
  • Conant,R. & Collins,J.T. 1991. A Field Guide to Reptiles and Amphibians of Eastern/Central North America, 3rd ed. Houghton Mifflin (Boston/New York), xx + 450 p.
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
  • Hahn D E 1979. Leptotyphlops humilis (Baird and Girard). Western blind snake. Catalogue of American Amphibians and Reptiles (232): 1-4 - get paper here
  • Heimes, P. 2016. Snakes of Mexico. Chimaira, Frankfurt, 572 pp
  • Jameson, David H.;Flury, Alvin G. 1949. The reptiles and amphibians of the Sierra Vieja Range of southwestern Texas. Texas Journal of Science 1 (2): 54-77 - get paper here
  • Klauber, Laurence M. 1939. A new subspecies of the western worm snake. Transactions of the San Diego Society of Natural History 9 (14): 67-68 - get paper here
  • Lazcano D, Nevárez-de los Reyes M, García-Padilla E, Johnson JD, Mata-Silva V, DeSantis DL, Wilson LD. 2019. The herpetofauna of Coahuila, Mexico: composition, distribution, and conservation status. Amphibian & Reptile Conservation 13(2) [General Section]: 31–94 (e189) - get paper here
  • Lemos-Espinal JA, Smith GR, Gadsden-Esparza H, Valdez-Lares R, Woolrich-Piña GA 2018. Amphibians and reptiles of the state of Durango, Mexico, with comparisons with adjoining states. ZooKeys 748: 65-87 - get paper here
  • Lemos-Espinal JA, Smith GR, Woolrich-Piña GA 2018. Amphibians and reptiles of the state of San Luis Potosí, Mexico, with comparisons with adjoining states. ZooKeys 753: 83-106 - get paper here
  • Lemos-Espinal, Julio A. (ed.) 2015. Amphibians and Reptiles of the US - Mexico Border States / Anfibios y Reptiles de los Estados de la Frontera México - Estados Unidos. Texas A&M University Press, College Station, Texas. x + 614 pp.; ISBN 978-1-62349-306-6. - get paper here
  • Lemos-Espinal, Julio A. and James R. Dixon 2013. Amphibians and Reptiles of San Luis Potosí. Eagle Mountain Publishing, xii + 300 pp.
  • Milstead, William W. 1953. Ecological distribution of the lizards of the La Mota Mountain Region of Trans-Pecos Texas. Texas Journal of Science 5 (4): 403-415 - get paper here
  • Nevárez-de-los-Reyes, Manuel, David Lazcano, Elí García-Padilla, Vicente Mata-Silva, Jerry D. Johnson and Larry David Wilson. 2016. The Herpetofauna of Nuevo León, Mexico: Composition, Distribution, and Conservation. Mesoamerican Herpetology 3 (3): 558–638 - get paper here
  • Tennant, A. & Bartlett, R.D. 2000. Snakes of North America - Eastern and Central Regions. Gulf Publishing, Houston, TX, 588 pp.
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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