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Rhagerhis moilensis (REUSS, 1834)

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Higher TaxaLamprophiidae, Psammophiinae, Colubroidea, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesE: Moila Snake
G: Moilanatter 
SynonymColuber moilensis REUSS 1834: 142
Coelopeltis producta GERVAIS 1857
Rhagerrhis moilensis — PETERS 1862
Rhagerrhis producta — SCLATER 1891:26
Coelopeltis producta — WERNER 1894: 85
Coelopeltis moilensis — BOULENGER 1896
Coelopeltis cordofanensis WERNER 1919: 506 (fide BÖHME & DE PURY 2011)
Malpolon moilensis — PARKER 1931: 522
Coelopeltis moilensis — ANGEL 1932: 386
Malpolon moilensis — HAAS 1957: 80
Rhamphiophis moilensis — KRAMER & SCHNURRENBERGER 1963
Malpolon moilensis — LEVITON & ANDERSON 1967: 182
Rhamphiophis maradiensis CHIRIO & INEICH 1991 (fide TRAPE & MANÉ 2006: 10)
Malpolon moilensis — MATTISON 1995: 221
Scutophis moilensis — BRANDSTÄTTER 1995
Malpolon moilensis — SCHLEICH, KÄSTLE & KABISCH 1996: 499
Scutophis moilensis — GENIEZ et al. 2004
Scutophis moilensis — SCHLÜTER 2005
Malpolon moilensis —TRAPE & MANÉ 2006
Scutophis moilensis — PADIAL 2006
Malopolon moilensis — MATTISON 2007: 245 (in error)
Malpolon moilensis — LARGEN & SPAWLS 2010: 548
Scutophis moilensis — BARATA et al. 2011
Rhagerhis moilensis — BÖHME & DE PURY 2011
Rhagerhis moilensis — WALLACH et al. 2014: 645
Rhagerhis maradiensis — WALLACH et al. 2014: 645 
DistributionMorocco, Western Sahara, Algeria, Tunisia, Libya, Egypt, Israel, Sinai, Jordan, Syria, Iraq, SW Iran, Saudi Arabia, Oman, United Arab Emirates (UAE), Kuwait,
Mauritania, Niger, Mali, Sudan, Eritrea, Ghana ?

Type locality: Moilah (= Al Muwaylih), Saudi Arabia.

Rhamphiophis maradiensis: Niger (Maradi); Type locality: “aux environs de Gari'n Bakwai, 50 km au sud-ouest de Maradi. au Niger”; Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesHolotype: SMF 20017, Syntype: ZMB 2635 (fide J. Hallermann, pers. comm.)
Holotype: MNHN 1989-2739, female [Rhamphiophis maradiensis]
Syntype: ZMB 10831 [Coelopeltis producta] 
CommentType speces: Coelopeltis producta GERVAIS 1857 is the type species of the genus Rhagerhis PETERS 1862: 274. For a diagnosis of the genus see BÖHME & DE PURY 2011.

Synonymy: TRAPE & MANÉ 2006 synonymized Rhamphiophis maradiensis CHIRIO & INEICH 1991 with Rhagerhis (Malpolon) moilensis. However, L. Chirio (pers. comm. 23 May 2013) disagrees, because “Malpolon (Scutophis) moilensis presents a caracteristic agressive behaviour, producing a flat neck a bit like a Naja. Then this species always shows 2 big dark spots on the edges of the neck. R. maradiensis doesn't show neither these spots, nor the agressive behaviour. It is a very timid snake, also thinner than M. moilensis.”

Type species: Coluber moilensis REUSS 1834 is the type species of Scutophis BRANDSTÄTTER 1995. Coelopeltis productus is the type species of the genus Rhagerhis PETERS 1862

Diagnosis (genus): (1) the skull structure (Szyndlar 1988) that results in the typical head shape of Moila snakes (see also Brandstätter 1996); (2) elongated neck ribs that are on average 3 mm longer than in equal-sized Malpolon (for details see de Pury 2010, and Fig. 2) and facilitate the spreading of the neck into a hood-shaped pattern (Fig. 1) characteristic of R. moilensis; (3) the dorsal scale microdermatoglyphic pattern that is drastically different in Rhagerhis moilensis as compared with both Mal­polon insignitus and M. monspessulanus (Böhme & Pury 2011). 
References
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