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Rhynchocalamus satunini (NIKOLSKY, 1899)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Palestine Kukri Snake
G: Schwarzkopf-Zwergnatter, Schwarzkopfnatter 
SynonymContia satunini NIKOLSKY 1899: 449
Oligodon melanocephalus septentrionalis WERNER 1905
Contia satunini — WERNER 1917
Oligodon melanocephalus satunini — CHERNOV 1937
Rhynchocalamus satunini — REED & MARX 1959
Rhynchocalamus melanocephalus satunini — DAREVSKY 1970
Rhynchocalamus melanocephalus satunini — FRANZEN & BISCHOFF 1995
Rhynchocalamus melanocephalus satunini — ANANJEVA et al. 2006: 182
Rhynchocalamus satunini — ŠMÍD et al. 2015 
DistributionE Turkey, S Armenia, Iraq (Kurdistan), W Iran, Azerbaijan

Type locality: "Prov. Elizabetpol in Transcaucasia, prope [near] flum. Arax" [found by K. Satunin in August 1893 in the vicinity of Megri [=Meghri], on the Axes River; now in Armenia].  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ZISP 9343 
Diagnosis 
CommentDistribution: not in Iraq fide Habeeb & Rastegar-Pouyani 2016; see map in ŠMÍD et al. 2015: 382. For a map see Sindaco et al. 2013. 
References
  • Al-Quran, S. 2009. The Herpetofauna of the Southern Jordan. American-Eurasian J. Agric. & Environ. Sci., 6 (4): 385-391 [this journal has a dubious record, see http://goo.gl/81SBPv]
  • Ananjeva, NB, Orlov NL; Khalikov RG; Darevsky IS; Ryabov IS & Barabanov AV 2006. The Reptiles of North Eurasia. Taxonomic Diversity, Distribution, Conservation Status [this comprises the territory of the former Soviet Union and Mongolia]. Pensoft Series Faunistica 47, 250 pp.
  • Arakelyan, Marine S.; Felix D. Danielyan, Claudia Corti, Roberto Sindaco, and Alan E. Leviton 2011. The Herpetofauna of Armenia and Nagorno-Karabakh. SSAR, Salt Lake City, 154 pp. [ISBN: 978-0-916984-84-7.]
  • AVCI, A. & ILGAZ, Ç. & KUMLUTAS, Y. & OLGUN, K. & BARAN, I. 2007. Morphology and distribution of Rhynchocalamus melanocephalus saturimi (NIKOLSKY, 1899) in Turkey. Herpetozoa 20: 82–85 - get paper here
  • Avci, A., Dinçaslan, Y.E., Ilgaz, Ç. & Üzüm, N. 2008. Contributions to the distribution and morphology of Rhynchocalamus melanocephalus melanocephalus (Jan 1862) (Reptilia, Colubridae) in Turkey. North-Western Journal of Zoology 4 (1): 161-166 - get paper here
  • Avcı, Aziz; Çetin Ilgaz, Mahdi Rajabizadeh, Can Yılmaz, Nazan Üzüm, Dominique Adriaens, Yusuf Kumlutaş, Kurtuluş Olgun 2015. Molecular Phylogeny and Micro CT-Scanning Revealed Extreme Cryptic Biodiversity in Kukri Snake, Muhtarophis gen. nov., a New Genus for Rhynchocalamus barani (Serpentes: Colubridae). Russ. J. Herpetol. 22 (3): 159-174
  • Baha el Din, S. 2006. A guide to the reptiles and amphibians of Egypt. The American University in Cairo Press, Cairo. xvi + 359 pp.
  • Bar, Aviad and Guy Haimovitch 2012. A Field Guide to Reptiles and Amphibians of Israel. Pazbar LTD, 246 pp. - get paper here
  • Berger-Dell'mour, H. 1986. Zur Herpetofauna des Golan [Syria]. Ann. Naturhist. Mus. in Wien (Ser. B) 87: 59-67. - get paper here
  • Boettger, O. [as O. Böttger] 1880. Die Reptilien und Amphibien von Syrien, Palaestina und Cypern. Ber. Senckenb. Naturforsch. Ges., Frankfurt/M., 1879-1880: 132-219 - get paper here
  • Corkill, N. L. and Cochrane, J. A. 1966. The snakes of the Arabian Peninsula and Socotra. J. Bombay nat. Hist. Soc. 62 (3): 475-506 (1965) - get paper here
  • Damhoureyeh, S.A.; Qarqaz, M.A.; Baker, M.A.; Himdan, N.; Eid, E. & Amr, Z.S. 2009. Reptiles and Amphibians in Dibbeen Nature Reserve, Jordan. Vertebrate Zoology 59 (2): 169 – 177
  • Darevsky, I. S. 1970. Systematic status of Rhynchocalamus melanocephalus satunini Nik. (Serpentes, Colubridae) previously included in the genus Oligodon. Zool. Zhurnal 49: 1685-1690.
  • Disi, A.M.; Modry, D.; Necas, P. & Rifai, L. 2001. Amphibians and reptiles of the Hashemite Kingdom of Jordan. Edition Chimaira, Frankfurt, 408 pp.
  • Egan, D. 2007. Snakes of Arabia. Motivate Publishing, Dubai, 208 pp.
  • Engelmann, W.E. et al. 1993. Lurche und Kriechtiere Europas. Neumann Verlag (Radebeul, Germany), 440 pp.
  • Esterbauer, H. 1985. Zur Herpetofauna Südwestsyriens. Herpetofauna 7 (38): 23-34 - get paper here
  • Franzen M; Bischoff W 1995. Erstnachweis von Rhynchocalamus melanocephalus melanocephalus für die Turkei. Salamandra 31 (2): 107-122 - get paper here
  • Gasperetti,J. 1988. Snakes of Arabia. Fauna of Saudi Arabia 9: 169-450
  • Geniez, Philippe 2018. Snakes of Europe, North Africa and the Middle East. Princeton University Press, 384 pp. - get paper here
  • Günther,A. 1864. Report on a collection of reptiles and fishes from Palestine. Proc. Zool. Soc. London 1864: 488-493 - get paper here
  • Hraoui-Bloquet, S.; Sadek, R.A.; Sindaco, R. & Venchi, A. 2002. The herpetofauna of Lebanon: new data on distribution. Zoology in the Middle East 27: 35-46 - get paper here
  • Jan, G. 1862. Enumerazione sistematico delle specie d'ofidi del gruppo Calamaridae. Arch. Zool., Anat. Fisiol., 2: 1-76. - get paper here
  • Jan, G. 1865. Iconographie générale des ophidiens. 13. Livraison. [Homalosoma mite]. J.B. Bailière et Fils, Paris - get paper here
  • Kamali, Kamran 2020. A Field Guide to the Reptiles and Amphibians of Iran. Edition Chimaira, Frankfurt am Main, Germany (www.chimaira.de). 574 pp.
  • Leviton,A.E.; Anderson,S.C.; Adler, K.; Minton,S.A. 1992. Handbook to Middle East Amphibians and Reptiles. SSAR, Oxford, Ohio (Contr. to Herpetol. No. 8), 1-252
  • Nikolsky, A. M. 1899. Contia satunini n. sp., and Agama ruderata Oliv., from Transcaucasia [in Russian]. Annuaire Musée Zoologique de l’Académie Impériale des Sciences de St.-Pétersbourg, 4: 449-451 - get paper here
  • Olgun, K.; Avci, A. Ilgaz, C.; Üzüm, N. & Yilmaz, C. 2007. A new species of Rhynchocalamus (Reptilia: Serpentes: Colubridae) from Turkey. Zootaxa 1399: 57–68 - get paper here
  • Rajabizadeh, M. 2018. Snakes of Iran. [in Farsi] Tehran: Iranshenasi, 496 pp. [2017 ?] - get paper here
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Reed, Charles A. & Marx, Hymen 1959. A herpetological collection from Northeastern Iraq. Transactions of the Kansas Academy of Science 62 (1): 91-122 - get paper here
  • Schmidt, Karl P. 1933. A new snake (Rhynchocalamus arabicus) from Arabia. Zoological Series of Field Museum of Natural History 20: 9-10 - get paper here
  • Schmidt,K.P. 1939. Reptiles and amphibians from Southwestern Asia. Publ. Field Mus. nat. Hist., zool. Ser., 24: 49-92 - get paper here
  • Sindaco, R.; Alberto Venchi & Cristina Grieco 2013. The Reptiles of the Western Palearctic, Volume 2: Annotated Checklist and Distributional Atlas of the Snakes of Europe, North Africa, Middle East and Central Asia, with an Update to Volume 1. Edizioni Belvedere, Latina (Italy), 543 pp. - get paper here
  • ŠMÍD, JIŘÍ; GABRIEL MARTÍNEZ, JURGEN GEBHART, JAVIER AZNAR, JAVIER GÁLLEGO, BAYRAM GÖÇMEN, PHILIP DE POUS, KARIN TAMAR & SALVADOR CARRANZA 2015. Phylogeny of the genus Rhynchocalamus (Reptilia; Colubridae) with a first record from the Sultanate of Oman. Zootaxa 4033 (3): 380–392 - get paper here
  • Tuniyev, B.S.; N.L. Orlov, N.B. Ananjeva and A.L. Aghasyan 2019. Snakes of the Caucasus: taxonomic diversity, distribution, conservation. St. Petersburg, Moscow. KMK Scientific Press. 2019. 276 pp. - get paper here
  • Werner, F. 1917. Reptilien aus Persien (Provinz Fars). Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 67:191—220. - get paper here
  • Werner,Y.L. 1988. Herpetofaunal survey of Israel (1950-1985), with comments on Sinai and Jordan and on zoogeographical heterogeneity. in: Yom-Tov,Y. & Tchernov, E. (eds.) The zoogeography of Israel, Dr. W. Junk Publishers, Dordrecht (Netherlands); ISBN 90-6193-650-0
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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