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Sitana devakai AMARASINGHE, INEICH & KARUNARATHNA, 2015

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards) 
Subspecies 
Common NamesE: Devaka’s Fan-throated Lizard
Sinhala (local): Devakagë Theli Katussa
French: Sitane de Devaka 
SynonymSitana devakai AMARASINGHE, INEICH & KARUNARATHNA in AMARASINGHE et al. 2015
Litana Ponticereana [sic] — KELAART (1854) [partim]
Sitana ponticeriana — SMITH (1935) [partim]
Sitana ponticeriana — TAYLOR (1957) [partim]
Sitana ponticeriana — WERMUTH (1967) [partim]
Sitana ponticeriana — MANAMENDRA-ARACHCHI & LIYANAGE (1994) [partim]
Sitana ponticeriana — ERDELEN (1998) [partim]
Sitana ponticeriana — DAS & DE SILVA (2005) [partim]
Sitana ponticeriana — DE SILVA (2006) [partim]
Sitana ponticeriana — SOMAWEERA & SOMAWEERA (2009) [partim]
Sitana ponticeriana — MANTHEY (2010) [partim]
Sitana ponticeriana ponticeriana — DERANIYAGALA (1953) [partim]
Sitana ponticertiana [sic]—BAHIR & SURASINGHE (2005) [partim] 
DistributionSri Lanka, India (Tamil Nadu)

Type locality: Nagagamuwa-Puttalam, Sri Lanka (8°10′N, 79°50′E), elevation 2 m  
Reproduction 
TypesHolotype. WHT 0111E, Male, SVL 45.6 mm, collected by K. Manamendra-Arachchi & D. Gabadage, 9 October 1993.
Paratypes. Males, WHT 0174A–B, respective SVL 41.1 mm and 40.0 mm, collected at Palavi-Puttalam, Sri Lanka; Male, WHT 1417, SVL 43.9 mm, collected at Angamuwewa-Rajanganaya, Sri Lanka; Male, WHT 1424A, SVL 39.9 mm, collected at Mundel-Puttalam, Sri Lanka; Female, WHT0174C, SVL 42.7 mm, collected at Palavi- Puttalam, Sri Lanka; Females, WHT 0111B–C, respective SVL 42.0 mm and 44.9 mm, collected at Nagagamuwa- Puttalam, Sri Lanka; Sub adult male, WHT 1424B, SVL 38.6 mm, collected at Mundel-Puttalam, Sri Lanka; Sub adult females, WHT 0111A,D, respective SVL 37.6 mm and 38.1 mm, collected at Nagagamuwa-Puttalam, Sri Lanka. All those specimens were collected by D. Gabadage. 
CommentDiagnosis (adult males). Sitana devakai sp. nov. differs from each congener by having the following characters: S. bahiri sp. nov.: longer throat-fan ADW 29.9–32.5% of AG (vs. shorter, ADW 28.3–29.2% of AG), midbody scale rows 57–59 (vs. 49–56), ventral scale number 100–108 (vs. 87–89), abdominal scales pointed (vs. bluntly pointed), equal and regular lateral scales with intermediate enlarged scales (vs. unequal and irregular lateral scales with intermediate enlarged scales), breeding-males with bright red colour patch (vs. bleached orange colour patch); S. deccanensis: smaller SVL 40.0–45.6 mm (vs. larger SVL above 60.0 mm), snout longer ES 57.2–63.2% of HW (vs. shorter, ES 53.1–53.4% of HW), throat-fan shorter ADW 29.9–32.5% of AG (vs. longer, ADW 50.3–60.2% of AG), femur shorter FEL 72.2–78.4% of TBL (vs. longer, FEL 79.7–82.9% of TBL), foot longer FOL 154.7–180.2 of HL (vs. shorter, FOL 131.9–144.6% of HL), supralabials 8–9 (vs. 12), midbody scale rows 57–59 (vs. 66–70), ventral scale number 100–108 (vs. 82–92), subdigital lamellae on toe III 15–16 (vs. 13), enlarged scales above the tympanum present (vs. absent), lateral scales with intermediate enlarged scales (vs. without enlarged scales), breeding-males with bright red colour patch restricted to mid throat-fan (vs. bright red patch extending to vent), single pale stripe running from the snout up to the shoulder (vs. 3–4 bright stripes from snout and behind the eye up to the shoulders); S. ponticeriana: femur shorter FEL 72.2–78.4% of TBL (vs. longer, FEL 80.0–84.8% of TBL), ventral scale number 100–108 (vs. 65–69), abdominal scales pointed (vs. rounded); Sitana sivalensis complex: throat-fan extending up to the chest (vs. not extending to the level of axilla), foot longer with FOL 154.7–180.2 of HL (vs. shorter, FOL 120.0–144.3% of HL). 
EtymologyThe species epithet is an eponym Latinised in the genitive singular, honouring Prof. Devaka Keerthi Weerakoon, for his remarkable contributions to biodiversity conservation in Sri Lanka. He worked as the Technical Advisor of the 2007 and 2012 Red Lists of Threatened Fauna and Flora in Sri Lanka and was responsible for their publication. As a consultant to the Department of Wildlife Conservation, he has provided technical support for the preparation and implementation of protected areas, produced management plans, managed alien invasive species programs in protected areas, mapped habitats, carried out protected area gap analyses programs and conducted bird surveys. His contributions to the ecology of Asian elephants and reduction of human-elephant conflicts are remarkable. 
References
  • AMARASINGHE, A. A. THASUN; IVAN INEICH, D. M. S. SURANJAN KARUNARATHNA,<br />W. MADHAVA S. BOTEJUE & PATRICK D. CAMPBELL 2015. Two new species of the genus Sitana Cuvier, 1829 (Reptilia: Agamidae) from Sri Lanka, including a taxonomic revision of the Indian Sitana species. Zootaxa 3915 (1): 067–098 - get paper here
 
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