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Smaug depressus (FITZSIMONS, 1930)

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Higher TaxaCordylidae (Cordylinae), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Flat Dragon Lizard, Zoutpansberg Girdled Lizard 
SynonymZonurus barbertonensis depressus FITZSIMONS 1930: 24
Zonurus laevigatus FITZSIMONS 1933
Cordylus laevigatus — FITZSIMONS 1943
Cordylus warreni depressus — FITZSIMONS 1943
Cordylus warreni laevigatus — LOVERIDGE 1944
Smaug warreni depressus — STANLEY et al. 2011
Smaug warreni depressus — BATES et al. 2014: 211
Smaug depressus — STANLEY & BATES 2014 
DistributionRepublic of South Africa (Middle and Eastern Soutpansberg [Limpopo] through the northern Krüger National Park and south to Duiwelskloof). Elevation up to 1500 m.

Type locality: Newgate, near Louis Trichardt, Northern Transvaal.  
ReproductionOvoviviparous. 
TypesHolotype: DNMNH (= TM) 4621 
Diagnosis 
CommentDistribution: see map in STANLEY & BATES 2014: Figure 2. 
EtymologyApparently named after the depressed head. 
References
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • FitzSimons, V. F. 1930. Descriptions of new South African Reptilia and Batrachia, with distribution records of allied species in the Transvaal Museum collection. Annals Transvaal Mus. 14: 20-48. - get paper here
  • FitzSimons, V.F. 1943. The lizards of South Africa. Transvaal Museum Memoir No.1 (Pretoria), 528 pp.
  • FitzSimons, VIVIAN 1933. Description of five new lizards from the Transvaal and Southern Rhodesia. Annals Transvaal Mus. 15 (2): 273-280. - get paper here
  • Loveridge,A. 1944. Revision of the African lizards of the family Cordylidae. Bull. Mus. comp. Zool. Harvard 95 (1): 1-118 - get paper here
  • Mouton, le Fras; Alexander Flemming; Michael Bates and Chris Broeckhoven 2018. The relationship between generation gland morphology and armour in Dragon Lizards (Smaug): a reassessment of ancestral states for the Cordylidae. Amphibia-Reptilia 39 (4): 445–456 - get paper here
  • Stanley, E. L. and Bates, M. F. 2014. Here be dragons: a phylogenetic and biogeographical study of the Smaug warreni species complex (Squamata: Cordylidae) in southern Africa. Zoological Journal of the Linnean Society 172 (4): 892–909; doi: 10.1111/zoj.12187 - get paper here
  • Stanley, Edward L.; Aaron M. Bauer; Todd R. Jackman, William R. Branch, P. Le Fras N. Mouton 2011. Between a rock and a hard polytomy: rapid radiation in the rupicolous girdled lizard (Squamata: Cordylidae). Molecular Phylogenetics and Evolution, 58(1): 53-70. - get paper here
 
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