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Stenocercus johaberfellneri KÖHLER & LEHR, 2015

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Higher TaxaTropiduridae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymStenocercus johaberfellneri KÖHLER & LEHR 2015
Stenocercus species — LEHR 2002: 87 
DistributionPeru (Huarmey)

Type locality: Malvas, -9.91683, -77.65133, 3200 m, Departamento Ancash, Provincia Huarmey, Peru  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: MUSM 20229, an adult male, collected 3 February 1997 by Edgar Lehr. Field tag number EL 99. Paratypes. 11 (5 males, 6 females). All from Malvas, Departamento Ancash, Provincia Huarmey, Peru, collected on 2–3 February 1997 by Edgar Lehr: MUSM 20225, -9.91833, -77.65800, 3200 m. MUSM 20226, - 9.92850, -77.64967, 3055 m. MUSM 20227, SMF 80231–33, -9.92850, -77.64517, 3120 m. MUSM 20228, SMF 80234, -9.93217, -77.65500, 3130 m. MUSM 20229–30, -9.91683, -77.65133, 3200 m. SMF 80235, -9.93217, - 77.65333, 3140 m. 
DiagnosisDiagnosis: Stenocercus johaberfellneri differs from all other congeners except S. latebrosus and S. ornatissimus by the combination of having (1) a well developed oblique neck fold with a deep mite pocket underneath; (2) a well developed antehumeral fold with a deep mite pocket underneath; (3) scales on posterior surface of thighs granular; (4) vertebral scales similar in size and shape to adjacent scale rows; (5) three caudal whorls per autotomic segment. Stenocercus johaberfellneri differs from S. latebrosus by having lateral nuchals less than half the size of dorsal nuchals (vs. lateral nuchals similar in size to dorsal nuchals in S. latebrosus). Stenocercus johaberfellneri differs from S. ornatissimus by having larger dorsal scales, 47–53, mean 50.0, vertebral scales (vs. dorsal scales smaller, 52–59, mean 55.4, vertebral scales in S. ornatissimus), 46–53, mean 49.2, scales around midbody (vs. 49–60, mean 52.6, in S. ornatissimus), the presence of a postfemoral pocket (Type 2 of Torres-Carvajal 2007) in adult males (vs. postfemoral pocket absent in S. ornatissimus), a throat pattern of bold black streaks or reticulations in adult males (vs. throat with small black spots or fine reticulations in S. ornatissimus), and a dusky or grayish venter without black spots or reticulations (vs. at least some black spots on pectoral region and anterior abdomen, often entire venter with small black spots in S. ornatissimus). 
CommentSympatry: Epictia alfredschmidti, Philodryas simonsii, and Sibynomorphus petersi (misidentified as S. vagus in Lehr 2002 and Lehr et al. 2002 as pointed out by Cadle 2007).

Abundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe name johaberfellneri is a patronym for Johannes Haber Fellner, Germany, in recognition of his support of taxonomic studies through the BIOPAT program. 
References
  • Cadle, J.E. 2007. The snake genus Sibynomorphus (Colubridae: Dipsadinae: Dipsadini) in Peru and Ecuador, with comments on the systematics of Dipsadini. Bull. Mus. Comp. Zool. Harvard 158(5):183-283 - get paper here
  • KÖHLER, GUNTHER & EDGAR LEHR 2015. Two new species of lizards of the genus Stenocercus (Iguania, Tropiduridae) from central Peru. Zootaxa 3956 (3): 413–427 - get paper here
  • Lehr, E. 2002. Amphibien und Reptilien in Peru. Natur und Tier-Verlag (Münster), 208 pp. - get paper here
  • Lehr, E.; Köhler, G. & Streit, B. 2002. Die Herpetofauna von Mittelperu entlang eines Transektes von der pazifischen Küste bis in die Hochanden (Amphibia et Reptilia). Faun. Abh. Mus. Tierk. Dresden 22 (2): 361-392
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Mendoza, A., Ramírez, C., Barrera, D., & Aguilar-Puntriano, C. 2021. A new species of the genus Stenocercus (Iguania: Tropiduridae) from the Peruvian Pacific coast (Ica region). Salamandra 57 (1): 1-14
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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