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Timon pater (LATASTE, 1880)

IUCN Red List - Timon pater - Least Concern, LC
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Higher TaxaLacertidae, Lacertinae, Sauria, Lacertoidea, Squamata (lizards)
Subspecies 
Common Names 
SynonymLacerta ocellata DAUDIN 1802 part.
Lacerta ocellata pater LATASTE 1880
Lacerta ocellata var. viridissima — BOETTGER 1885: 466
Lacerta ocellata var. pater — BOULENGER 1920
Lacerta ocellata — LAURENT 1935: 346
Lacerta ocellata pater — HEDIGER 1935
Lacerta lepida pater — ANGEL 1946
Lacerta (Gallotia) lepida — PETERS 1961
Lacerta lepida pater — LANGERWERF 1981
Lacerta pater — BISCHOFF 1982
Lacerta pater — SCHLEICH, KÄSTLE & KABISCH 1996: 412
Timon pater — MAYER & BISCHOFF 1996: 169
Timon pater — ARNOLD et al. 2007
Timon pater — AHMADZADEH et al. 2016 
DistributionAlgeria, Tunisia, Morocco ?, elevation up to 2111 m (Atlas, Algeria)

pater: eastern part of Morocco, E of Oued Moulouya Valley, Algeria

Type locality: not specifically given by Lataste, who says “ " Tres commun a Alger, Sétif, Bone, Batna, Lambèze, etc...". Mayer 2015 effectively restricted the type locality to “Sétif in Algerien” although he did not explicitly mentioning a restriction.  
Reproductionoviparous 
Typessyntype: MCZ R-4613 
Diagnosis 
CommentSubspecies: Timon pater tangitana (BOULENGER, 1889) has been elevated to full species status.

Distribution: See AHMADZADEH et al. 2016: Fig. 1 for a map. Not in Morocco but possibly in Morocco (Martínez et al. 2019, and pers. comm. 18 Jan 2021).

Elevation: the max. elevation previously reported was 2650 m, but this represented t. tangitanus, not T. pater (Bouam et al. 2017). 
References
  • Ahmadzadeh, Faraham; Morris Flecks, Miguel A. Carretero, Wolfgang Böhme, Flora Ihlow, Paschalia Kapli, Andreia Miraldo and Dennis Rödder 2016. Separate histories in both sides of the Mediterranean: phylogeny and niche evolution of ocellated lizards. Journal of Biogeography; DOI: 10.1111/jbi.12703 - get paper here
  • Arnold, E.N.; Arribas, O. & Carranza, S. 2007. Systematics of the Palaearctic and Oriental lizard tribe Lacertini (Squamata: Lacertidae: Lacertinae), with descriptions of eight new genera. Zootaxa 1430: 1-86 - get paper here
  • Bezaz, Y. I., Hadjab, R., Khammar, H., Redjaimia, L., & Saheb, M. 2021. First data on the diversity of the herpetofauna of the Oum El Bouaghi Region (Northeast of Algeria). Ecol. Env. & Cons. 27 (3): 983-989
  • Bischoff, W. 1985. Lacerta, (Timon) pater LATASTE, 1880. Sauria 7 (4) Suppl.: 029–030 - get paper here
  • Boettger, O. 1885. Liste der von Hrn. Dr. med. W. Kobelt in Algerien und Tunisien gesammelten Kriechtiere. In W. Kobelt (editor), Reiseerinnerungen aus Algerien und Tunis. Berichte der Senckenbergischen Naturforschungs Gesellschaft 1885: 452–475 - get paper here
  • Bouam, Idriss; Abdeldjabar Necer, Messaoud Saoudi 2017. Highest altitudinal record of the ocellated lizard Timon pater (Lataste, 1880) (Squamata: Lacertidae). Herpetology Notes 10: 101-102 - get paper here
  • Boulenger, George A. 1891. Catalogue of the reptiles and batrachians of Barbary (Morocco, Algeria, Tunisia), based chiefly upon the notes and collections made in 1880-1884 by M. Fernand Lataste. Tr. Zool. Soc. 13: 93-164
  • Busack S D. 1987. Morphological and biochemical differentiation in Spanish and Moroccan populations of the lizard, Lacerta lepida. Journal of Herpetology 21 (4): 277-284 - get paper here
  • Escoriza, Daniel 2017. Patterns of alpha diversity among Tunisian lizards (Lacertidae). Journal of Arid Environments - get paper here
  • Galán, F. 1931. Batracios y reptiles del Marruecos español. Bol. Soc. Española Hist. Nat. 31(5): 361-367 - get paper here
  • Hediger, H. 1935. Herpetologische Beobachtungen in Marokko. Verhandlungen der Naturforschenden Gesellschaft in Basel 46: 1-49
  • Langerwerf, B. 1981. Nigrinos bei der Nachzucht von Lacerta lepida pater. Herpetofauna 3 (12): 21-22 - get paper here
  • Lataste, F. 1880. Diagnoses de reptiles nouveaux d’Algérie. Le Naturaliste 1:299, 306—307, 325 - get paper here
  • Laurent, P. 1935. Contribution à la connalssance de la faune des Vertebres du Maroc (Batraciens, Reptiles, Mammiferes). Bull. Soc. d’Hist. nat. de l’Afrique du Nord 26: 344-359
  • Martínez del Mármol, Gabriel; D. James Harris, Philippe Geniez, Philip de Pous, and Daniele Salvi 2019. Amphibians and Reptiles of Morocco. Edition Chimaira, Frankfurt am Main, Germany, 478 pp - get paper here
  • Mateo, J. A., A. Santana & P. Geniez 2004. Moroccan ocellated lizard in Western Sahara; answer to an old enigma. Herpetozoa 17 (3/4): 187-189 [2004] - get paper here
  • Mateo, J.A.;Lopezjurado, L.F.;Guillaume, C.P. 1996. Proteic and morphological variations in ocellated lizards (Lacertidae): A complex of species across the Strait of Gibraltar. Comptes Rendus de L Academie Des Sciences Serie III - Sciences de La Vie - Life Sciences 319 (8): 737-746
  • Mayer, W., and Bischoff, W. 1996. Beiträge zur taxonomischen Revision der Gattung Lacerta (Reptilia: Lacertidae). Teil 1: Zootoca, Omanosaura, Timon und Teira als eigenständige Gattungen. Salamandra 32 (3): 163-170 - get paper here
  • Mediani, Mohamed; José Carlos Brito, Soumia Fahd 2015. Atlas of the amphibians and reptiles of northern Morocco: updated distribution and patterns of habitat selection. Basic and Applied Herpetology 29: - get paper here
  • Odierna,G. et al. 1990. Karyological differences beween Lacerta lepida and Lacerta pater. Journal of Herpetology 24 (1): 97-99 - get paper here
  • Peters, G. 1961. Die Perleidechse (Lacerta lepida Daudin) gehört zum Subgenus Gallotia Boulenger. Mitteilungen aus dem Zoologischen Museum in Berlin, 37(2): 272–285.
  • Rouag, Rachid; Berrahma, Ilhem; Luiselli, Luca 2006. Food habits and daily activity patterns of the North African ocellated lizard Timon pater from northeastern Algeria. Journal of Natural History 40 (21-22): 1369-1379 - get paper here
  • Saoudi, Messaoud; Abdeldjabar Necer, Idriss Bouam, Farouk Khelfaoui & Oussama Saadi 2017. Daily activity, biometry and diet of the North African ocellated lizard Timon pater in Mount Chélia, north-eastern Algeria. African Journal of Herpetology 66 (1): 1-11 - get paper here
  • Schleich, H.H., Kästle,W., Kabisch, K. 1996. Amphibians and Reptiles of North Africa. Koeltz, Koenigstein, 627 pp.
  • Schlüter, U. 2004. Die Herpetofauna des Mittleren Atlas von Marokko. Reptilia (Münster) 9 (45): 42-49 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
 
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