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Tretanorhinus variabilis DUMÉRIL, BIBRON & DUMÉRIL, 1854

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
SubspeciesTretanorhinus variabilis variabilis DUMÉRIL, BIBRON & DUMÉRIL 1854
Tretanorhinus variabilis binghami SCHWARTZ & OGREN 1956
Tretanorhinus variabilis insulaepinorum BARBOUR 1916
Tretanorhinus variabilis lewisi GRANT 1941
Tretanorhinus variabilis wagleri JAN 1863 
Common NamesE: Caribbean Water Snake
G: Karibische Wasserschlange 
SynonymTretanorhinus variabilis DUMÉRIL, BIBRON & DUMÉRIL 1854: 349
Tretanorhinus variabilis — BOULENGER 1893: 282
Helicops variabilis — GARMAN 1884: 33
Tretanorhinus variabilis — SCHWARTZ & HENDERSON 1991: 628
Tretanorhinus variabilis — WALLACH et al. 2014: 736

Tretanorhinus variabilis binghami SCHWARTZ and OGREN 1956
Tretanorhinus variabilis binghami — SCHWARTZ & HENDERSON 1988

Tretanorhinus variabilis insulaepinorum BARBOUR 1916
Tretanorhinus insulae-pinorum BARBOUR 1916
Tretanorhinus variabilis insulaepinorum — WOOD 1939
Tretanorhinus variabilis insulaepinorum — SCHWARTZ & HENDERSON 1988

Tretanorhinus variabilis lewisi GRANT 1941
Tretanorhinus variabilis lewisi GRANT 1941: 46
Tretanorhinus variabilis lewisi — SCHWARTZ & HENDERSON 1988

Tretanorhinus variabilis variabilis DUMÉRIL, BIBRON & DUMÉRIL 1854
Tretanorhinus variabilis var. adnexus BOCOURT 1891
Tretanorhinus variabilis var. cubanus BOCOURT 1895 (nom. subst. pro adnexus)
Tretanorhinus variabilis variabilis — GRANT 1946: 104
Tretanorhinus gaigeae GRANT 1946: 104
Tretanorhinus variabilis variabilis — SCHWARTZ & HENDERSON 1988

Tretanorhinus variabilis wagleri JAN 1863
Helicops wagleri JAN 1863: 248
Tretanorhinus variabilis wagleri — GRANT 1946: 103
Tretanorhinus variabilis wagleri — SCHWARTZ & HENDERSON 1988 
DistributionCuba, Isla de la Juventud, Cayman I

variabilis: Cuba, from La Habana Prov. in the west, east into Guantánamo Prov. (except for region about Sierra Maestra, Granma Prov.). Type locality: Unknown.

binghami: Cuba, Granma Prov., lowlands and lower foothilîs of Sierra Maestra. Type locality: Finca Bucares, 22 km S Bueycito, in the Rio Yao, Granma Province, Cuba.

insulaepinorum: Isla de la Juventud; Type locality: Isla de la Juventud.

lewisi: Cayman Is.: Grand Cayman I.; Type locality: North Side, Grand Cayman Island, Cayman Islands.

wagleri: Cuba, Pinar del Rio Province. Type locality: Brasil (in error)  
Reproductionoviparous 
TypesSyntypes: MNHN 7161, MNHN 7346.
Holotype: MNHN 7349 [adnexus]
Holotype: CAS-SU 14440 [gaigeae]
Holotype: NCSM 84101 (was: ChM 55.1.61 = Charleston Museum) [binghami]
Holotype: MCZ 44890 [lewisi]
Holotype: CM 311 [insulaepinorum]
Holotype: unlocated [wagleri] 
Diagnosis 
CommentSynonymy: Grant 1946 suspected that T. insulaepinorum is a synonym of T. wagleri.

For illustrations see Barbour and Ramsden 1919, Neill 1965, Petzold 1967, Petzold 1969.

Type species: Tretanorhinus variabilis DUMÉRIL, BIBRON & DUMÉRIL 1854: 349 is the type species of the genus Tretanorhinus DUMÉRIL, BIBRON & DUMÉRIL 1854.

Habitat: T. v. gaigae is “probably confined to brackish water” (Grant 1946: 117). 
Etymology 
References
  • Barbour, T. 1916. The reptiles and amphibians of the Isle of Pines. Annals of the Carnegie Museum 10 (1-2): 297-308. - get paper here
  • Barbour, T. and C. T. Ramsden. 1919. The herpetology of Cuba. Mem. Mus. Comp. Zool., Harvard Coll. 47: 69-213. - get paper here
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Duméril, A.M.C., G. BIBRON & A.H.A. DUMÉRIL 1854. Erpétologie générale ou Histoire Naturelle complète des Reptiles. Vol. 7 (partie 1). Paris, xvi + 780 S. - get paper here
  • Garman,Samuel 1884. The reptiles and batrachians of North America. Mem. Mus. comp. Zool, Cambridge (Massachusetts), 8 (3): xxxiv + 185 pp. [1883] [CNAH reprint 10] - get paper here
  • Grant, C. 1946. Notes on Tretanorhinus of Cuba and the Isle of Pines. J. Dept. Agric. Puerto Rico 30 (2): 102-117
  • Neill, W.T. 1965. Notes on aquatic snakes, Natrix and Tretanorhi nus, in Cuba. Herpetologica 21 (1): 62-67. - get paper here
  • Petzold, H.G. 1967. Some remarks on the breeding biology and the keeping of Tretanorhinus variabilis, a water snake of Cuba. Herpetologica 23(3): 242-46. - get paper here
  • Petzold, H.G. 1969. Zur Haltung and Fortpflanzungsbiologieeiniger kubanischer Schlangen im Tierpark Berlin. Salamandra 5:124-40 - get paper here
  • Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96 - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Schwartz, A. and Ogren, L.H. 1956. A collection of reptiles and amphibians from Cuba, with the descriptionof two new forms. Herpetologica 12 (2): 91-110 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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