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Trimeresurus toba DAVID, PETRI, VOGEL & DORIA, 2009

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Higher TaxaViperidae, Crotalinae, Colubroidea, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesToba pitviper. 
SynonymTrimeresurus (Popeia) toba DAVID, PETRI, VOGEL & DORIA 2009
Lachesis gramineus — BAUMANN 1913: 272 (nec Coluber gramineus SHAW 1802)
Trimeresurus cf. sabahi — VOGEL et al. 2004: 52
Popeia sabahi — SANDERS et al. 2006: 355 (nec Trimeresurus popeiorum sabahi REGENASS & KRAMER 1981)
Trimeresurus toba — DAVID et al. 2009
Trimeresurus (Popeia) toba — DAVID et al. 2011
Popeia toba — WALLACH et al. 2014: 576 
DistributionIndonesia (Sumatra)

Type locality: “si rambé”, now sirambi, about 13 km se of Balige, off the southern shore of Danau Toba (Indonesia, Sumatra island, province of Sumatera Utara.  
Reproduction 
TypesHolotype: msng 30988, adult female; collected by elio modigliani, 1891. 
CommentDiagnosis. A species of the genus Trimeresurus Lacépède, 1804, characterized by (1) hemipenes long, reaching in situ at least the 25th sC, without spines; (2) 1st supralabial distinct from nasal; (3) 21 msr; (4) overall green coloration in males and females without darker crossbands; (5) absence of a postocular streak in both males and females; (6) thin, white ventrolateral stripe present in males, faint but present or absent in females; (7) rather short tail in females with a ratio taL/tL between 0.148 and 0.157; (8) occipital scales smooth or weakly keeled; and (9) temporal scales large, as large as posterior temporals. 
Etymologythe specific epithet is the name of the volcanic toba massif in which the type locality is located; the same name is applied to the large Lake toba. it is a noun in apposition. 
References
  • David, P., Petri, M., Vogel, G. & G. Doria 2009. A new species of pitviper of the genus Trimeresurus (Popeia) from northern Sumatra (REPTILIA, SQUAMATA, VIPERIDAE). Annali del Museo Civico di Storia Naturale "G. Doria", Genova 100: 323-346
  • DAVID, PATRICK; GERNOT VOGEL & ALAIN DUBOIS 2011. On the need to follow rigorously the Rules of the Code for the subsequent designation of a nucleospecies (type species) for a nominal genus which lacked one: the case of the nominal genus Trimeresurus Lacépède, 1804 (Reptilia: Squamata: Viperidae). Zootaxa 2992: 1–51 - get paper here
  • Sumontha,M., Kunya,K., S. G. Pauwels,O., Nitikul,A., and Punnadee,S. 2011. Trimeresurus (Popeia) phuketensis, a New Pitviper (Squamata: Viperidae) from Phuket Island, Southwestern Thailand. Russ. J. Herpetol. 18 (3): 11-17 - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, 1237 pp.
 
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