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Tropidolaemus wagleri (BOIE, 1827)

IUCN Red List - Tropidolaemus wagleri - Least Concern, LC
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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Wagler's Palm Viper, Wagler's Pit Viper, Temple pitviper
G: Waglers Bambusotter 
SynonymCophias wagleri BOIE in SCHLEGEL 1826 (nomen nudum)
Cophias wagleri BOIE 1827: 561
[Tropidolaemus] wagleri WAGLER 1830
Trigonocephalus wagleri — SCHLEGEL 1837
Trimesurus maculatus GRAY 1842
Trimesurus maculatus GRAY 1842
Trimesurus sumatranus — GRAY 1842 (nec Coluber sumatranus RAFFLES 1822)
Trigonocephalus sumatranus var. — CANTOR 1847
Trigonocephalus wagleri, var. sumatrenis GRAY 1849
Trigonocephalus formosus — GRAY 1849 (nec Trigonocephalus formosus MÜLLER & SCHLEGEL 1842)
Tropidolaemus wagleri — DUMÉRIL et al. 1854: 1524
Trimeresurus sumatranus — GÜNTHER 1858
Tropidolaemus maculatus — PETERS 1859
Tropidolaemus sumatranus — PETERS 1862
Trimeresurus wagleri — GÜNTHER 1864
Trimeresurus subannulatus var. immaculatus PETERS 1872: 42
Bothrops wagleri — MÜLLER 1882
Lachesis wagleri — BOULENGER 1896
Lachesis wagleri — BOULENGER's var. D — VOLZ 1904
Lachesis wagleri — BOULENGER's var. A — COHN 1905
Lachesis wagleri var. B — HOLTZINGER-TENEVER 1917
Trimeresurus wagleri wagleri — TAYLOR 1922
Lachesis sumatranus — OUWENS 1916
Lachesis wagleri — DE ROOIJ 1917: 286
Trimeresurus (Tropidolaemus) wagleri — CAMPBELL & BRODIE Jr. 1992
Tropidolaemus wagleri — WELCH 1994: 117
Tropidolaemus wagleri — MANTHEY & GROSSMANN 1997: 411
Tropidolaemus wagleri — COX et al. 1998: 23
Tropidolaemus wagleri — MCDIARMID, CAMPBELL & TOURÉ 1999: 349
Tropidolaemus wagleri — TU et al. 2000
Tropidolaemus wagleri — WALLACH et al. 2014: 748 
DistributionS Vietnam ? (Minh Hai and Song Be) (Orlov et al., 2003),
S Thailand (Phang Nga, Pattani, Surat Thani, Nakhon Si Tammarat, Narathiwat, and Yala (Nabhitabhata et al., 2004).
West Malaysia (incl. Pulau Pangkor), Singapore, Indonesia (Sumatra, Bangka, Mentawei Archipelago, Natuna Islands, Nias, and Riau Archipelago)

Type locality: “Asien”; corrected by neotype designation to “Deli: rivière de Bedagneh (Est de Sumatra)”, now Bedagai River (about 3°30’N, 99°13’E), Sumatera Barat Province, Sumatra, Indonesia.  
Reproductionovovivparous 
TypesNeotype: MNHN-RA 1879.0708, adult female, collected by Mr. Lucien Rochet (designated by Vogel et al. 2007) 
DiagnosisDiagnosis: A species of the genus Tropidolaemus, characterized by (1) internasals always in contact; (2) a strong ontogenetic variation of the background body colour: black (never green) in adult females, whereas males and juveniles retain a vividly green background colour; (3) a strong ontogenetic variation of the pattern: yellow crossbands around the body in adult females, white spots in adult and juvenile males, white crossbars in juvenile females; (4) a black postocular stripe in adult females and a white and red one in males and juveniles of both sexes; (5) a banded belly in adult females and a uniform belly in males and juveniles; (6) 21–23 dorsal scale rows at midbody (DSR) at midbody in males and 23–27 in females, usually feebly keeled in males and distinctly keeled in females; (7) 143–152 ventrals (VEN) in males and 134–147 VEN in females; (8) 50–55 subcaudals (SC) in males and 45–54 in females (from VOGEL et al. 2007). 
CommentVenomous!

Nomenclature: For a discussion of nomenclatorial issues see David & Vogel, 1996, McDiarmid et al. (1999: 350) and Savagae et al. 2012. These authors argue that the author should be BOIE 1827 instead of WAGLER 1830 because BOIE doesn’t describe the species but refers to a plate in SEBA (1735) which is valid (Art. 12b and 72b of the Code). The ICZN has ruled that Trigonocephalus [Cophias] wagleri was established by Schlegel (1826) as a separate taxon and not as a replacement name (ICZN 2014, Opinion 2350)

Illustration: Portrait in Elaphe 5 (4): 81.

Distribution: Reported from Sulawesi by MÜLLER (1895: 865) but actually subannulatus and laticinctus. Reports from Vietnam are questionable as these specimens were obtained from a local dealer. Not on Borneo fide Stuebing et al. 2014.

Type Species: Cophias wagleri F. Boie, 1827 is the type species of the genus Tropidolaemus WAGLER 1830. 
EtymologyNamed after Johann Georg Wagler (1800-1832), German physician and zoologist. See Michahelles in Wagler 1833 for a short obituary. Apparently, Wagler accidentally shot himself while hunting, and died later from his wounds. 
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