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Tympanocryptis argillosa MELVILLE, CHAPLIN, HIPSLEY, SARRE, SUMNER & HUTCHINSON, 2019

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Higher TaxaAgamidae (Amphibolurinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Claypan Earless Dragon 
SynonymTympanocryptis argillosa MELVILLE, CHAPLIN, HIPSLEY, SARRE, SUMNER & HUTCHINSON 2019: 23 
DistributionAustralia (South Australia)

Type locality: E Side of Lake Hope Channel,12 km SSE Red Lake Yard, Strzelecki Desert, South Australia, 28°19’1100 S, 139°13’1400 E.  
Reproduction 
TypesHolotype. SAMA R49360, adult female. Collected by M.H. on 9 April 1997.
Paratypes. NMV D3119, Lake Eyre, South Australia, 28°22’ S, 137°22’ E, adult female; SAMA R3619, Lake Palankarinna, South Australia, 28°4600 S, 138°2500 E, adult female; R46073, Artemia Point, Hunt Peninsula, Lake Eyre North, South Australia, 28°4100 S, 137°2200 E, adult male; R51039, 6.3 km NNW of Wimma Hill, South Australia, 27°17’3200 S, 140°17’2600 E, adult male; R51360–61, 4.4 km NNW of Beal Hill, Simpson Desert Regional Reserve, South Australia, 26°35’4300 S, 137°49’5100 E, adult male; R54088, 4.1 km SW of Kalamurina homestead, South Australia, 27°55’3600 S, 137°57’2200 E, adult male: R54247, 23.5 km W of Kannakaninna Waterhole, Kalamurina Station, South Australia, 27°53’4000 S, 137°44’2300 E, adult female; R54248, 23.5 km W of Kannakaninna Waterhole, Kalamurina Station, South Australia, 27°53’4000 S, 137°44’2300 E, adult male. 
DiagnosisDiagnosis: A species of Tympanocryptis with rostral scale separated from the canthus rostralis, nasal scale extending dorsally across the canthus and bordered below by enlarged scales, well-developed lateral neck fold, no lateral skin fold, weakly keeled dorsal head scales, five-lined pattern present but light lines usually discontinuous, ventral surface white. 
CommentHabitat. Specimens mostly associated with the clayey swales between the dunes of the sandy deserts where it occurs.

Distribution: see map in Melville et al. 2019: 6 (Fig. 1: species B) 
EtymologyAdjectival form of the Latin word for clay, argilla, alluding to the preference of this species for the heavier soiled clay swales that are commonly encountered between the dunes of the Strzelecki and southern Simpson Deserts. 
References
  • Melville J, Chaplin K, Hipsley CA, Sarre SD, Sumner J, Hutchinson M. 2019. Integrating phylogeography and high-resolution X-ray CT reveals five new cryptic species and multiple hybrid zones among Australian earless dragons. R. Soc. open sci. 6: 191166 - get paper here
 
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