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Tympanocryptis pseudopsephos DOUGHTY, KEALLEY, SHOO & MELVILLE, 2015

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Higher TaxaAgamidae (Amphibolurinae), Sauria, Iguania, Squamata (lizards) 
Subspecies 
Common NamesGoldfields pebble-mimic dragon 
SynonymTympanocryptis pseudopsephos DOUGHTY, KEALLEY, SHOO & MELVILLE 2015 
DistributionAustralia (Western Australia: Goldfields region)

Type locality: 31 km south-west of Doolgunna Homestead (25°52'33"S, 119°01'41"E), Western Australia  
Reproduction 
TypesHolotype: WAM R154790, an adult male collected on 5 April 2004 by B. Maryan and B. Budrey (Fig. 14). Paratypes. WAM R114550 (female), 67 km south of Capricorn Roadhouse, 18 km east of Bulloo Down homestead (24°00'S, 119°45'E); WAM R136459 (female), Norseman Area (32°12'S, 121°47'E); WAM R164347 (male), 46.1 km east of Leonora (28°51'36"S, 121°48'12"E); WAM R164350 (female), 11.8 km east of Leonora (28°54'23"S, 121°26'44"E); WAM R167478 (male), 49.5 km north of Leinster (27°22'36"S, 120°35'10"E). 
DiagnosisDiagnosis. Distinguished from other Tympanocryptis by the following combination of character states: rotund body shape, presence of two pre-cloacal pores, lack of longitudinal stripes on the dorsum, presence of enlarged scales with raised spines arranged in short transverse rows of 4–7 scales, concave snout formed by protruding mouth, scales on snout rugose with feeble keels, rostral width ~1–2 times height, keels on scales of upper arm not aligned, well-defined row of enlarged scales at anterior and dorsal edge of thigh forming conspicuous ridge, scales on dorsal surface of thigh not aligned, ventrals smooth, and reddish-brown ground color, lacking lines but often possessing a dark charcoal-like wash over the head and dorsum.

Comparisons with other species. Based on its widespread distribution, we provide comparisons of T. diabolicus sp. nov. to all species treated here except T. cephalus. Tympanocryptis pseudopsephos sp. nov. is distinguished from T. gigas by smaller body size, more rotund body shape, concave (versus convex) snout, possessing enlarged scales on dorsum arranged in short transverse rows (versus slightly enlarged scales scattered on dorsum), conspicuous ridge on front of thigh formed by enlarged row of scales, and scales on top of thigh homogeneous.
Tympanocryptis pseudopsephos sp. nov. is distinguished from T. diabolicus sp. nov. by possessing enlarged dorsal scales in short transverse rows in rows of 4–7 scales (versus 2–5), snout concave (versus straight or convex), rostral scale width 2 times height (versus 3 times), keels of scales on upper arm not aligned, scales on upper thigh not aligned, and ventrals smooth or slightly raised (versus with low keels).
Tympanocryptis pseudopsephos sp. nov. is distinguished from T. fortescuensis sp. nov. by possessing enlarged dorsal scales in short transverse rows in rows of 4–7 scales (versus 2–5), concave snout (versus straight to convex), scales on snout rugose with feeble keels (versus with low keels), rostral scale 3 times wider than high (versus 2 times), keels on dorsal surface of upper arm not aligned, keels of scales on top of thigh not aligned (versus usually aligned), ventrals smooth or slightly raised (versus with low keels), and reddish-brown color overlain with black (versus light brown). 
CommentHabitat. Recorded from Mallee and Mulga woodlands, often with stones strewn on the ground. 
Etymologypseudopsephos means ‘false pebble’, with pseudo Latin for ‘false’ and psephos is Greek for ‘pebble’. The name is an allusion to the pebble-mimicking abilities of this species. 
References
  • DOUGHTY, PAUL; LUKE KEALLEY, LUKE P. SHOO & JANE MELVILLE 2015. Revision of the Western Australian pebble-mimic dragon species-group (Tympanocryptis cephalus: Reptilia: Agamidae) Zootaxa 4039 (1): 085–117 - get paper here
 
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