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Tytthoscincus ishaki GRISMER, 2006

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Higher TaxaScincidae, Sphenomorphinae, Scincoidea, Sauria, Squamata (lizards) 
Subspecies 
Common NamesTioman Island Forest Skink 
SynonymSphenomorphus ishaki GRISMER 2006
Sphenomorphus ishaki — GRISMER 2011
Tytthoscincus ishaki — GRISMER et al. 2016 
DistributionWest Malaysia (Pulau Tioman, Pahang)

Type locality: at an altitude of 850 m elevation on Gunung Kajang, Pulau Tioman, Pahang, West Malaysia, in hill dipterocarp forest.  
Reproduction 
TypesHolotype: ZRC 2.6157, adult female, collected on 20 March 2004 by P.L. Wood, Jr. beneath a rotten log. 
CommentDiagnosis: Sphenomorphus ishaki can be distinguished from all other Sundaland species of Sphenomorphus by the unique combination of having an adult SVL of 38–41 mm; 30–32 smooth, midbody scales; 68–73 paravertebral scales; 60–70 ventral scales; four supraoculars; parietals contacting supraoculars; prefrontals in contact; two enlarged posterior superciliary scales projecting dorsomedially; two loreal scales; six supralabial scales; five infralabial scales; 11 keeled, subdigital lamellae beneath the fourth toe; two enlarged, preanal scales; adpressed limbs not overlapping; body bands absent; faint, dark, dorsolateral stripe; distinct, white, postorbital stripe [from GRISMER 2006] 
EtymologyNamed after Muhamad Ishak Mat Sohor of Kampung Juara, Pulau Tioman who has served GRISMER as “guide and companion on many treks up Gunung Kajang and who has taught him a wealth of information about the natural history of the forest of Pulau Tioman.” 
References
  • Grismer, L. Lee 2006. Two new species of skinks (Genus Sphenomorphus Fitzinger 1843) from the Seribuat Archipelago, West Malaysia. Herpetological Natural History 9 (2): 151-162.
  • Grismer, L. Lee. 2008. A new species of insular skink (Genus Sphenomorphus Fitzinger 1843) from the Langkawi Archipelago, Kedah, West Malaysia with the first report of the herpetofauna of Pulau Singa Besar and an updated checklist of the herpetofauna of Pulau Langkawi. Zootaxa 1691:53-66 - get paper here
  • GRISMER, L. LEE; MOHD ABDUL MUIN, PERRY L. WOOD, JR., SHAHRUL ANUAR, & CHARLES W. LINKEM 2016. The transfer of two clades of Malaysian Sphenomorphus Fitzinger (Squamata: Scincidae) into the genus Tytthoscincus Linkem, Diesmos, & Brown and the description of a new Malaysian swamp-dwelling species Zootaxa 4092 (2): 231–242 - get paper here
  • GRISMER, L. LEE; PERRY L. WOOD, JR., EVAN S.H. QUAH, SHAHRUL ANUAR, EHWAN B. NGADI, NUR AMALINA MOHD IZAM and NORHAYATI AHMAD 2017. Systematics, ecomorphology, cryptic speciation and biogeography of the lizard genus Tytthoscincus Linkem, Diesmos & Brown (Squamata: Scincidae) from the sky- island archipelago of Peninsular Malaysia. Zoological Journal of the Linnean Society XX: 1–37 - get paper here
  • Grismer, L.L. 2007. A new species of small montane forest floor skink (genus Sphenomorphus Fitzinger 1843) form Southern peninsular Malaysia. Herpetologica 63 (4): 544-551 - get paper here
  • Grismer, L.L. 2011. Amphibians and reptiles of the Seribuat Archipelago. Edition Chimaira, Frankfurt, 239 pp.
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Grismer, L.L.; Ahmad, N. & Kin Onn, C. 2009. A new, diminutive, upland Sphenomorphus Fitzinger 1843 (Squamata: Scincidae) from the Belum-Temengor Forest Complex, Peninsular Malaysia. Zootaxa 2312: 27–38 - get paper here
  • Grismer, L.L.; Wood Jr., P.L. & Grismer, J.L. 2009. A New Insular Species of Skink of the Genus Sphenomorphus Strauch 1887 (Squamata: Scincidae) from Pulau Perhentian Besar, Terengganu, Peninsular Malaysia. Tropical Life Sciences Research, 20(1): 51–69
 
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