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Tytthoscincus perhentianensis (GRISMER, WOOD & GRISMER, 2009)

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Higher TaxaScincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Perhentian Island Forest Skink 
SynonymSphenomorphus perhentianensis GRISMER, WOOD & GRISMER 2009
Sphenomorphus perhentianensis — GRISMER 2011
Tytthoscincus perhentianensis — GRISMER et al. 2016 
DistributionWest Malaysia (Pulau Perhentian Besar)

Type locality: Pulau Perhentian Besar on the trail behind Water Colours Resort (05°54.054 N, 102°44.343 E), Terengganu, Peninsular Malaysia  
Reproduction 
TypesHolotype: LSUHC 8075, adult male, collected on October 18, 2007 by Perry L. Wood Jr.. 
Diagnosis 
CommentSphenomorphus perhentianensis is differentiated from all other species of Sundaland and Malay Peninsula Sphenomorphus because it is small SVL (30.0mm) and having 29 midbody scale rows, smooth as opposed to striated dorsal scales, 65 paravertebrals, 61 ventrals, four supraoculars, parietals contacting the posterior-most supraocular, one medially projecting superciliary scale, two loreals, six supralabials and infralabials, ten lamellae beneath the fourth toe, smooth subdigital lamellae, enlarged preanal scales, no body bands, a dark brown, diffuse, dorsolateral stripe extending to just past the axilla, a cream coloured dorsolateral stripe on the nape and anterior-most portion of the body, and no cream coloured postorbital stripe. Table 2 shows the distribution of these character states across all 36 species )from GRISMER et al. 2009].

Abundance: only known from the type specimen (Meiri et al. 2017). 
Etymologynamed after the type locality. 
References
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • GRISMER, L. LEE; MOHD ABDUL MUIN, PERRY L. WOOD, JR., SHAHRUL ANUAR, & CHARLES W. LINKEM 2016. The transfer of two clades of Malaysian Sphenomorphus Fitzinger (Squamata: Scincidae) into the genus Tytthoscincus Linkem, Diesmos, & Brown and the description of a new Malaysian swamp-dwelling species. Zootaxa 4092 (2): 231–242 - get paper here
  • GRISMER, L. LEE; PERRY L. JR. WOOD, AMIRRUDIN B. AHMAD, B. S. BAIZUL-HAFSYAM, M. AFIQ-SHUHAIMI, SYED A. RIZAL, EVAN S. H. QUAH 2018. Two new Tytthoscincus Linkem, Diesmos, & Brown (Squamata; Scincidae) from Peninsular Malaysia and another case of microsyntopy between ecologically specialized, unrelated, leaf-litter species. Zootaxa 4425 (1): 087–107 - get paper here
  • GRISMER, L. LEE; PERRY L. WOOD, JR., EVAN S.H. QUAH, SHAHRUL ANUAR, EHWAN B. NGADI, NUR AMALINA MOHD IZAM and NORHAYATI AHMAD 2017. Systematics, ecomorphology, cryptic speciation and biogeography of the lizard genus Tytthoscincus Linkem, Diesmos & Brown (Squamata: Scincidae) from the sky- island archipelago of Peninsular Malaysia. Zoological Journal of the Linnean Society XX: 1–37 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Grismer, L.L.; Ahmad, N. & Kin Onn, C. 2009. A new, diminutive, upland Sphenomorphus Fitzinger 1843 (Squamata: Scincidae) from the Belum-Temengor Forest Complex, Peninsular Malaysia. Zootaxa 2312: 27–38 - get paper here
  • Grismer, L.L.; Wood Jr., P.L. & Grismer, J.L. 2009. A New Insular Species of Skink of the Genus Sphenomorphus Strauch 1887 (Squamata: Scincidae) from Pulau Perhentian Besar, Terengganu, Peninsular Malaysia. Tropical Life Sciences Research, 20(1): 51–69 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
 
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