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Woodworthia maculata (GRAY, 1845)

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Higher TaxaDiplodactylidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Common NamesRaukawa gecko 
SynonymNaultinus maculatus GRAY 1845: 273 (non Hoplodactylus maculatus BOULENGER 1885)
Naultinus brevidactylus GRAY 1845: 273
Naultinus granulatus GRAY 1845: 273
Naultinus pacificus — BULLER 1871 (partim)
Naultinus granulatus — BULLER 1871
Hoplodactylus maculatus — BOULENGER 1885: 171
Hoplodactylus granulatus — BOULENGER 1885: 174
Woodworthia digitata GARMAN 1901: 4
Hoplodactylus pacificus — WERNER 1901
Dactylocnemis maculatus — HUTTON & DRUMMOND 1905
Gehyra pacifica — MERTENS 1934
Naultinus granulatus — CHRAPLIWY et al. 1961
Hoplodactylus digitatus — WERMUTH 1965
Hoplodactylus granulatus — WERMUTH 1965: 95 (partim)
Hoplodactylus maculatus — RÖSLER 2000: 90
Woodworthia maculatus — JEWELL 2008: 52
Woodworthia maculatus — NIELSEN et al. 2011
Woodworthia maculata — HITCHMOUGH et al. in CHAPPLE 2016 
DistributionNew Zealand (North Island and northern South Island, including offshore islands)

Type locality: New Zealand
Type locality: New Zealand [brevidactylus]
Type locality: New Zealand [granulatus]  
TypesLectotype: BMNH 1946.9.8.14, New Zealand, designated by Bauer (1990). 
CommentSynonymy: Thomas (1981) synonymizes Hoplodactylus maculatus with H. granulatus. CHRAPLIWY et al. (1961) synonymized N. brevidactylus and N. maculatus with H. granulatus.

Type species: Woodworthia digitata Garman 1901 = Woodworthia maculatus (Gray, 1845) is the type species of the genus Woodworthia GARMAN 1901. Hitchmough et al. in Chapple (2016: 102) pointed out that Woodworthia is feminine, so all species names should be feminine too.

Definition (genus). Pupils with crenulated margins; subdigital pads >2/3 of toe length; 7–15 lamellae under digit IV of pes; ventral skin translucent; mouth lining unpigmented, tongue with or without pigmented tip; precloacal pores of males continuous with femoral pores on under surface of thighs; SVL 38–85 mm [NIELSEN et al. 2011] 
EtymologyNamed after Mr. W. McM. Woodworth (1864–1912), assistant in charge of Vermes (worms) as well as Agassiz’ personal assistant in the running of the museum and a controversial figure in Boston of that era. Woodworthia is feminine fide Hitchmough et al. 2016. 
  • BELL, TRENT & AARON M. BAUER 2017. Predation on a free-ranging Raukawa gecko (Woodworthia maculata) by a purple rock crab (Leptograpsus variegatus) BioGecko (4): 20-25
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Buller, W. 1871. A list of the lizards inhabiting New Zealand, with descriptions. Transactions and Proceedings of the New Zealand Institute, 3 [1870]:4-11 - get paper here
  • Chrapliwy, P. S., H. M. Smith, and C. Grant. 1961. Systematic status of the geckonid lizard genera Gehyra, Peropus, Hoplodactylus and Naultinus. Herpetologica 17: 5-12 - get paper here
  • Cree, Alison and Kelly M. Hare 2010. Equal thermal opportunity does not result in equal gestation length in a cool-climate skink and gecko. Herp. Cons. Biol. 5 (2) - get paper here
  • Cree, Alison and Kelly M. Hare 2016. Reproduction and Life History of New Zealand Lizards. in: Chapple, D.G. (ed). New Zealand Lizards. Springer, pp. 169-206 - get paper here
  • Daugherty, Charles H.; Patterson, G. B.; Hitchmough, R. A. 1994. Taxonomic and conservation review of New Zealand herpetofauna. New Zealand Journal of Zoology 21 (4): 317-323
  • Freeman, Alistair 1993. Size differences between populations of Hoplodactylus maculatus in Canterbury, New Zealand. Herpetofauna (Sydney) 23 (1): 9-15
  • Garman, S. 1901. Some reptiles and batrachians from Australasia. Bull. Mus. Comp. Zool. Harvard 39: 1-14 - get paper here
  • Gaulke, M. & Altenbach, A. 2003. 100% pure New Zealand. Reptilia (Münster) 8 (41): 56-63 - get paper here
  • Gray, J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of die British Museum/Edward Newman, London: xxvii + 289 pp. - get paper here
  • Henle, K. 1981. Die Herpetofauna Neuseelands, Teil 4: Endemische Geckos. Herpetofauna 3 (15): 16-19 - get paper here
  • Hitchmough, Rodney A.; Geoffrey B. Patterson, and David G. Chapple 2016. Putting a Name to Diversity: Taxonomy of the New Zealand Lizard Fauna in: Chapple, D.G. (ed). New Zealand Lizards. Springer, pp. 87-108 - get paper here
  • Hoare, Joanne M. and Kelly M. Hare. 2006. Hoplodactylus maculatus Toxin consumption. Herpetological Review 37 (1): 86-87 - get paper here
  • Jewell, Tony 2008. A Photographic Guide to Reptiles and Amphibians of New Zealand [with corrections and comments in Chapple & Hitchmough 2009]. New Holland Publishers (NZ) Ltd,Auckland, 143 pp.
  • Macredie, Grant 1984. The lizard fauna of Castle Rock, Coromandel, New Zealand. Herpetofauna (Sydney) 15 (1-2): 37-40
  • Nielsen, Stuart V.; Aaron M. Bauer, Todd R. Jackman, Rod A. Hitchmough and Charles H. Daugherty 2011. New Zealand geckos (Diplodactylidae): Cryptic diversity in a post-Gondwanan lineage with trans-Tasman affinities. Molecular Phylogenetics and Evolution 59 (1): 1-22 - get paper here
  • Robb,J. 1980. New Zealand Amphibians and Reptiles in Colour. Collins, Auckland, 128 pp.
  • Rock, J. & Cree, A. 2003. Intraspecific variation in the effect of temperature on pregnancy in the viviparous gecko Hoplodactylus maculatus. Herpetologica 59 (1): 8-22 - get paper here
  • Rösler, H. 2000. Studien an den Begattungsorganen der Geckos (Reptilia: Gekkota) - 3. Die Hemipenismorphologie von Arten der Gattungen Hoplodactylus FITZINGER 1843, Naultinus GRAY 1842, Oedura GRAY 1842, Rhacodactylus FITZINGER 1843 und Strophurus FITZINGER 1843 (Gek Gekkota 2: 220-248
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Rösler, Herbert 2017. Gecko-Chorologie (Squamata: Gekkota). Gekkota (4): 1-160
  • Thomas, B.W. 1981. Hoplodactylus rakiurae n. sp. (Reptilia: Gekkonidae) from Stewart Island, New Zealand and comments on the taxonomic status of Heteropholis nebulosus McCann. New Zealand Journal of Zoology 8: 33- 47
  • van Winkel, D., Baling, M. & Hitchmough, R. 2018. Reptiles and Amphibians of New Zealand: A field guide. Auckland University Press, Auckland, 376 pp
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