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Anolis beckeri BOULENGER, 1881

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Higher TaxaDactyloidae, Iguania, Sauria, Squamata (lizards)
Common NamesE: Becker’s Lichen Anole
S: Abaniquillo Liquenado de Becker 
SynonymAnolis beckeri BOULENGER 1881
Anolis beckeri — BOULENGER 1885: 46
Anolis beckeri — GÜNTHER 1885: 45
Anolis beckeri — BARBOUR & COLE 1906: 149
Anolis pentaprion — STUART 1937 (part.)
Anolis pentaprion beckeri — ETHERIDGE 1959
Anolis pentaprion beckeri — STUART 1963
Anolis pentaprion beckeri — LINER 2007
Anolis pentaprion beckeri — LINER & CASAS-ANDREU 2008: 43
Anolis beckeri — KÖHLER 2010
Norops beckeri — NICHOLSON et al. 2012
Norops beckeri — MCCRANIE 2015
Norops beckeri — NICHOLSON et al. 2018 
DistributionMexico (Campeche, Chiapas, Quintana Roo, Tabasco, Veracruz, Oaxaca), Belize, Guatemala, Honduras, N Nicaragua

Type locality: “Yucatán”  
TypesHolotype: IRSNB 2010 
DiagnosisDiagnosis. A medium-sized species (SVL in largest specimen examined 60.6 mm in males, 55.4 mm in females) of the genus Anolis (sensu Poe, 2004) that differs from all Central American anoles except A. charlesmyersi, A. cristifer, A. fungosus, A. pentaprion, A. salvini, and A. utilensis by having (1) extremely short hind legs (fourth toe of adpressed hindlimb usually reaching to level of tympanum; ratio shank length / head length 0.49–0.84); (2) smooth, juxtaposed ventral scales; (3) a relatively short tail (ratio tail length / SVL 1.0–1.5). Anolis beckeri differs from A. charlesmyersi by having a pink dewlap in life with purple pigment between rows of gorgetal scales and regular rows of numerous small narrowly spaced gorgetal scales (versus a brick red dewlap with few large widely spaced gorgetal scales scattered across the dewlap). Anolis beckeri differs from A. cristifer by having a double row of slightly enlarged middorsal scales, not forming a serrated crest (versus a serrated middorsal crest, at least on posterior portion of dorsum, present in A. cristifer). Anolis beckeri differs from A. fungosus by lacking bony parietal protuberances (versus having a pair of small bony parietal protuberances posterior and lateral to interparietal plate). Anolis beckeri differs from A. pentaprion by lacking a pronounced serrated caudal crest (versus having a pronounced serrated caudal crest) and by having gorgetal rows with 4–10 widely spaced scales (versus gorgetal rows with 17–25 narrowly spaced scales). Anolis beckeri differs from A. salvini by having smooth, obliquely conical ventral scales (versus keeled), lacking a dark interorbital bar (versus present), and lacking enlarged postcloacal scales (versus males with a pair of greatly enlarged postcloacal scales). Anolis beckeri differs from A. utilensis by having the proximal subdigital scales of toes differentiated as slightly broadened lamellae (versus those scales granular). 
CommentSpecies group: Norops auratus Species Group (fide Nicholson et al. 2012).Belongs to the pentaprion group which includes seven species: fungosus, ibague, ortonii, pentaprion, sulcifrons, utilensis, and vociferans. Norops pentaprion cristifer is listed here as valid species.

Synonymy: McCranie (2009), Townsend and Wilson (2010a), and Wilson and Johnson (2010) implicitly regarded this name as a synonym of Norops pentaprion, but Köhler (2010) resurrected it as a distinct species. Poe considers the diagnostic characters suggested by Köhler as insufficient to distinguish between A. beckeri and utilensis (Poe 2016).

Distribution: not in Yucatan state (Mexico) according to GONZÁLEZ-SÁNCHEZ et al. 2017. 
EtymologyThe name beckeri is a patronym honoring the Belgian arachnolo- gist Leo ́ n Becker. 
  • Aguilar-López JL, Luría-Manzano R, Pineda E, Canseco-Márquez L 2021. Selva Zoque, Mexico: an important Mesoamerican tropical region for reptile species diversity and conservation. ZooKeys 1054: 127-153 - get paper here
  • Barbour, T., and L. J. Cole. 1906. Vertebrata from Yucatan. Reptilia, Amphibia and Pisces. Bull. Mus. Comp. Zool. Harvard 50: 146-159 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Boulenger, George A. 1881. Description of a new species of Anolis from Yucatan. Proc. Zool. Soc. London 1881: 921-922 - get paper here
  • Campbell, J.A. 1998. Amphibians and reptiles of northern Guatemala, the Yucatán, and Belize. Norman: University of Oklahoma Press, xiii + 380 pp. - get paper here
  • Castañeda-Hernández, Carlos, Luis Canseco-Márquez and Manuel Ernesto Vargas-Orrego. 2015. Additional distributional records for the state of Oaxaca, Mexico. Mesoamerican Herpetology 2(3): 368–370 - get paper here
  • Gray, Levi. 2015. Book Review: The Anoles of Honduras. Herpetological Review 46 (4): 655-657 - get paper here
  • Günther, A. C. L. G. 1885. Reptilia and Batrachia. Biologia Centrali-Américana. Taylor, & Francis, London, 326 pp. [published in parts from 1885-1902; reprint by the SSAR 1987] - get paper here
  • Johnson, Jerry D.; Vicente Mata-Silva, Elí García Padilla, and Larry David Wilson 2015. The Herpetofauna of Chiapas, Mexico: composition, distribution, and conservation. Mesoamerican Herpetology 2 (3): 272–329. - get paper here
  • Köhler, G. 2010. A revision of the Central American species related to Anolis pentaprion with the resurrection of A. beckeri and the description of a new species (Squamata: Polychrotidae). Zootaxa 2354: 1–18 - get paper here
  • Liner, Ernest A. 2007. A CHECKLIST OF THE AMPHIBIANS AND REPTILES OF MEXICO. Louisiana State University Occasional Papers of the Museum of Natural Science 80: 1-60 - get paper here
  • Liner, Ernest A., and Gustavo Casas-Andreu. 2008. Standard Spanish, English and Scientific Names of the Amphibians and Reptiles of Mexico. Herpetological Circular 38: 167 p.
  • McCranie, James R. 2015. A checklist of the amphibians and reptiles of Honduras, with additions, comments on taxonomy, some recent taxonomic decisions, and areas of further studies needed. Zootaxa 3931 (3): 352–386 - get paper here
  • McCranie, James R. 2018. The Lizards, Crocodiles, and Turtles of Honduras. Systematics, Distribution, and Conservation. Bulletin of the Museum of Comparative Zoology, Special Publication Series (2): 1- 666 - get paper here
  • McCranie, James R. and Gunther Köhler 2015. The Anoles (Reptilia: Squamata: Dactyloidae: Anolis: Norops) of Honduras. Systematics, Distribution, and Conservation. Bull. Mus. Comp. Zool. Harvard (Special Publication Series, No. 1): 1-280 [review in Q. J. Biol. 91: 227] - get paper here
  • NICHOLSON, KIRSTEN E. & GUNTHER KÖHLER 2014. A new species of the genus Norops from Darién, Panama, with comments on N. sulcifrons (Cope 1899) (Reptilia, Squamata, Dactyloidae). Zootaxa 3895 (2): 225–237 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Owens, John. B., Kimberley C. Carter, Cornelio Santiago Chablé Tesucún and Rowland Griffin. 2016. Reproductive and parental care notes for Norops beckeri (Boulanger, 1891) in northern Guatemala. Mesoamerican Herpetology 3(4): 1007–1010 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Poe, S. 2016. Book review: The Anoles of Honduras: Systematics, Dis­ tribution, and Conservation. Q. Rev. Biol. 91: 227 - get paper here
  • Solís, J. M., L. D. Wilson, and J. H. Townsend. 2014. An updated list of the amphibians and reptiles of Honduras, with comments on their nomenclature. Mesoamerican Herpetology 1: 123–144 - get paper here
  • Stuart,L.C. 1963. A checklist of the herpetofauna of Guatemala. Misc. Publ. Mus. Zool., Univ. Michigan (No. 122): 1-150 - get paper here
  • Sunyer, Javier 2014. An updated checklist of the amphibians and reptiles of Nicaragua. Mesoamerican Herpetology 1 (2): 186–202. - get paper here
  • Townsend, J. H., L. D. Wilson, M. Medina-Flores, E. Aguilar-Urbina, B. K. Atkinson et al. 2012. A premontane hotspot for herpetological endemism on the windward side of Refugio de Vida Silvestre Texíguat, Honduras. Salamandra 48 (2): 92-114 - get paper here
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